Publications


# Graduate student; ^ Undergraduate student

Journal Articles, Book Chapters, Proceedings, & Books

In press

147. Maxwell EC#, Koltz A^, Stamn J#, Beatty C^, Boudreaux C^, Carbonare J^, Clasper C^, Harden L^, Harris J^, Lawmaster K^, Lee J^, Polk J^, Polk J^, Yeager R^, da Costa Arantes I, Ohmer MEB, & Niemiller ML. Rediscovery of the Tennessee Cave Salamander (Gyrinophilus palleucus) McCrady, 1954 at Shelta Cave, Madison County, Alabama, USA, a former subterranean hotspot cave. Southeastern Naturalist.

146. Niemiller ML, Anderson CL#, Maxwell EC#, Midgette H, Quering C^, Richards M#, Wilson K, & Wirwa N. A noteworthy range extension of the federally endangered Alabama Cave Shrimp (Palaemonias alabamae) south of the Tennessee River in northern Alabama. Southeastern Naturalist.

2025

145. Benito JB#, Ober KA, Philips TK, Orndorff W, & Niemiller ML. Phylogenomics and biogeography of North American trechine cave beetles (Coleoptera: Carabidae) inferred from ultraconserved elements. Evolution 79: 1622–1642. https://doi.org/10.1093/evolut/qpaf103

144. Hart PB, Rincon-Sandoval M, Melendez-Vazquez F, Armbruster JW, Troyer EM, Bierstein OM#, Gough BJ, Betancur-R R, Niemiller ML, & Arcila D. Ancient climate changes and relaxed selection shape cave colonization in North American cavefishes. Proceedings of the Royal Society B, Biological Sciences 292: 20242516. https://doi.org/10.1098/rspb.2024.2516

143. Niemiller ML, Zigler KS, Curtis AN, Trapeni CM^, Slay ME, Culver DC, Hutchins BT, & Niemiller KDK. Out of sight and out of mind? The conservation status of subterranean biodiversity in the United States and Canada. Biodiversity and Conservation 34: 2851–2882. https://doi.org/10.1007/s10531-025-03099-6

142. Niemiller ML, Hart PM, Pirro S, & Arcila D. The complete genome sequences of 12 species of Percopsiformes. Biodiversity Genomes. https://doi.org/10.56179/001c.133607

2024

141. Kennedy KA^, Zigler KS, Cramphorn B#, Harden CW#, Helf K, Lewis JJ, Malabad TE, Milne MA, Niemiller ML, & Stephen CDR. Remarkably low genetic diversity in the widespread cave spider Phanetta subterranea (Araneae, Linyphiidae). Subterranean Biology 50: 105–118. https://doi.org/10.3897/subtbiol.50.135200

140. Cucalón RV#, Corush JB, Niemiller ML, Curtis AN, Hart PB, Kuhajda BR, Thomas MR, Metzke B, Davis MA, & Tan M. 2024. Population genomics and mitochondrial DNA reveal cryptic diversity in North American spring cavefishes (Amblyopsidae, Forbesichthys). Conservation Genetics 25: 1283–1301. https://doi.org/10.1007/s10592-024-01640-8

139. Costley TL#, Hopkins SR, Meng S^, Gajewski Z, & Niemiller ML. Summer cave use by Tricolored Bats declined in response to white-nose syndrome despite persistence in winter hibernacula in the southeastern United States. Journal of Mammalogy 105: 1032–1043. https://doi.org/10.1093/jmammal/gyae064

138. Reynolds RG, Comsa LC#, & Niemiller ML. Status and observations of the critically endangered Turks Island Skink (Spondylurus turksae). Reptiles & Amphibians 31: e22449. https://doi.org/10.17161/randa.v31i1.22449

137. Wilson KL#, Niemiller KDK, & Niemiller ML. The reproductive biology of the northern slimy salamander (Plethodon glutinosus) from a cave in northern Alabama, USA with notes on diet. Herpetological Conservation and Biology 19: 222–235. http://www.herpconbio.org/Volume_19/Issue_2/Wilson_etal_2024.pdf

136. Espinasa L, Gutierrez A^, Hinkle A, & Niemiller ML. A new genus and species of nicoletiid silverfish (Insecta: Zygentoma: Nicoletiidae) from caves of northern Alabama, USA. Subterranean Biology 49: 1–17. https://doi.org/10.3897/subtbiol.49.119986

135. Benito J#, Porter ML, & Niemiller ML. Comparative mitogenomic analysis of subterranean and surface amphipods (Crustacea, Amphipoda) with special reference to the family Crangonyctidae. BMC Genomics 25: 298. https://doi.org/10.1186/s12864-024-10111-w

134. Niemiller ML, Cannizzaro AG#, Sawicki TR, & Culver DC. A new species of Stygobromus Cope, 1872 (Amphipoda: Crangonyctidae) from a hypotelminorheic seepage spring in Washington, D.C., USA. Subterranean Biology 48: 117–146. https://doi.org/10.3897/subtbiol.48.112984

133. Tobin BW, Miller BV, Niemiller ML, & Erhardt AM. Expanding karst groundwater tracing techniques: incorporating population genetic and isotopic data to enhance flow-path characterization. Hydrology 11: 23. https://doi.org/10.3390/hydrology11020023

132. Lunghi E, Niemiller ML, & Bilandzija H. Editorial: Adaptations to subterranean environments. Frontiers in Ecology and Evolution 11: 1354954. https://doi.org/10.3389/fevo.2023.1354954

131. LaFrance BJ, Ray AM, Fisher RN, Grant EHC, Shaffer C, Beamer DA, Spear SF, Pierson TW, Davenport JM, Niemiller ML, Pyron RA, Glorioso BM, Barichivich WJ, Halstead BJ, Roberts KG, & Hossack BR. A dataset of amphibian species in U.S. national parks. Scientific Data 11: 32. https://doi.org/10.1038/s41597-023-02836-2

130. Saccò M, Mammola S, Altermatt F, Alther R, Bolpagni R, Brancelj A, Brankovits D, Fišer C, Gerovasileiou V, Griebler C, Guareschi S, Hose GC, Korbel K, Lictevout E, Malard F, Martínez A, Niemiller ML, Robertson A, Tanalgo KC, Bichuette ME, Borko S, Brad T, Campbell MA, Cardoso P, Celico F, Cooper SJB, Culver D, Di Lorenzo T, Galassi DMP, Guzik MT, Hartland A, Humphreys WF, Ferreira RL, Lunghi E, Nizzoli D, Perina G, Raghavan R, Richards Z, Reboleira ASPS, Rohde MM, Fernández DS, Schmidt SI, van der Heyde M, Weaver L, White NE, Zagmajster M, Hogg I, Gagnon MM, Allentoft ME, & Reinecke R. Groundwater is a globally threatened keystone ecosystem. Global Change Biology 30: e17066. https://doi.org/10.1111/gcb.17066

129. Hart PB, Niemiller ML, Armbruster JW, & Chakrabarty P. Population genomics of a cavefish species complex with implications for conservation. Conservation Genetics 25: 165–177. https://doi.org/10.1007/s10592-023-01562-x

2023

128. Niemiller ML, Zigler KS, Hinkle A, Stephen CDR, Cramphorn B#, Higgs J#, Mann N, Miller BT, Niemiller KDK, Smallwood K, & Hardy J. The Crystal-Wonder Cave System: a new hotspot of subterranean biodiversity in the southern Cumberland Plateau of south-central Tennessee, USA. Diversity 15: 801. https://doi.org/10.3390/d15070801

127. Niemiller ML, Slay ME, Inebnit T, Miller B, Tobin B, Cramphorn B#, Hinkle A, Jones BD, Mann N, Niemiller KDK, & Pitts S. Fern Cave: a hotspot of subterranean biodiversity in the Interior Low Plateau karst region of Alabama in the southeastern United States. Diversity 15: 633. https://doi.org/10.3390/d15050633

126. Mouser JB#, Johnston J, Niemiller ML, & Brewer SK. A fisherman’s tale: an unusual observation of the Ozark cavefish, Troglichthys (=Amblyopsis) rosae (Eigenmann). Southeastern Naturalist 22: N28–N33. https://doi.org/10.1656/058.022.0114

125. Harper LR, Niemiller ML, Benito JB#, Molano-Flores B, & Davis MA. BeeDNA: microfluidic environmental DNA metabarcoding as a tool for connecting plant and pollinator communities. Environmental DNA 5: 191–211. https://doi.org/10.1002/edn3.370

124. Griebler C, Hahn HJ, Mammola S, Niemiller ML, Weaver L, Sacco M, Bichuette ME, & Hose GC. Legal frameworks for the conservation and sustainable management of groundwater ecosystems. Pp. 551–571 in: Groundwater Ecology and Evolution (Malard F, Griebler C, & Retaux S, eds.). Academic Press.

123. Boulton AJ, Bichuette ME, Korbel K, Stoch F, Niemiller ML, Hose GC, & Linke S. Recent concepts and approaches for conserving groundwater biodiversity. Pp. 520–550 in: Groundwater Ecology and Evolution (Malard F, Griebler C, & Retaux S, eds.). Academic Press.

122. Venarsky M, Niemiller ML, Fišer C, Saclier N, & Moldovan OT. Life histories in groundwater organisms. Pp. 439–456 in: Groundwater Ecology and Evolution (Malard F, Griebler C, & Retaux S, eds.). Academic Press.

121. Zagmajster M, Ferreira RL, Humphreys WF, Niemiller ML, & Malard F. Patterns and determinants of richness and composition of the groundwater fauna. Pp. 141–164 in: Groundwater Ecology and Evolution (Malard F, Griebler C, & Retaux S, eds.). Academic Press.

2022

120. Mouser JB#, Brewer SK, Niemiller ML, Mollenhauer R, & Van Den Bussche RA. Lithology and disturbance drive cavefish and cave crayfish occurrence in the Ozark Highlands ecoregion, USA. Scientific Reports 12: 19559. https://doi.org/10.1038/s41598-022-21791-3

119. Grant EHC, Mulder KP, Brand AB, Chambers DB, Wynn A, Capshaw G#, Niemiller ML, Phillips JG, Kuchta SR, & Bell RC. Speciation with gene flow in a narrow endemic West Virginia cave salamander (Gyrinophilus subterraneus). Conservation Genetics 23: 727–744. https://doi.org/10.1007/s10592-022-01445-7

118. Niemiller ML, Hardman R, Thames D, Istvanko D, Davis MA, Ogle C, Niemiller KDK, Dooley KE, & Clark TM. The distribution and conservation status of the green salamander (Aneides aeneus) in Tennessee, USA. Herpetological Conservation and Biology 17: 249–265.

117. Mammola S, Meierhofer M, Borges P, Colado R, Culver D, Deharveng L, Delic T, Di Lorenzo T, Dražina T, Ferreira R, Fiasca B, Fiser C, Galassi D, Garzoli L, Gerovasileiou V, Griebler C, Halse S, Howarth F, Isaia M, Johnson J, Komerički A, Martínez A, Milano F, Moldovan O, Nanni V, Nicolosi G, Niemiller M, Pallarés S, Pavlek M, Piano E, Pipan T, Sánchez Fernández D, Santangeli A, Schmidt S, Wynne J, Zagmajster M, Zakšek V, & Cardoso P. Toward evidence-based conservation of subterranean ecosystems. Biological Reviews 97: 1476–1510. https://doi.org/10.1111/brv.12851

116. Niemiller ML, Davis MA, Tan M, Apodaca JJ, Dooley KE#, Cucalón RV, Benito J#, Niemiller KDK, Harden R, Thames D, & Istvanko D. Mitochondrial DNA and population genomics reveal additional cryptic diversity in the green salamander (subgenus Castaneides) species complex. Frontiers in Conservation Science 3: 890859. https://doi.org/10.3389/fcosc.2022.890859

115. Dooley KE#, Niemiller KDK, Sturm N#, & Niemiller ML. Rediscovery and phylogenetic analysis of the Shelta Cave crayfish (Orconectes sheltae Cooper & Cooper, 1997), a decapod (Decapoda, Cambaridae) endemic to Shelta Cave in northern Alabama, USA. Subterranean Biology 43: 11–31. https://doi.org/10.3897/subtbiol.43.79993

114. Ferreira RL, Bernard E, Júnior FW da C, Piló LB, Calux A, Souza-Silva M, Barlow J, Pompeu PS, Cardoso P, Mammola S, García AM, Jeffery WR, Shear W, Medellín RA, Wynne JJ, Borges PAV, Kamimura Y, Pipan T, Hajna NZ, Sendra A, Peck S, Onac BP, Culver DC, Hoch H, Flot J-F, Stoch F, Pavlek M, Niemiller ML, Manchi S, Deharveng L, Fenolio D, Calaforra J-M, Yager J, Griebler C, Nader FH, Humphreys WF, Hughes AC, Fenton B, Forti P, Sauro F, Veni G, Frumkin A, Gavish-Regev E, Fišer C, Trontelj P, Zagmajster M, Delic T, Galassi DMP, Vaccarelli I, Komnenov M, Gainett G, Tavares V da C, Kováč Ľ, Miller AZ, Yoshizawa K, Lorenzo TD, Moldovan OT, Sánchez-Fernández D, Moutaouakil S, Howarth F, Bilandžija H, Dražina T, Kuharić N, Butorac V, Lienhard C, Cooper SJB, Eme D, Strauss AM, Saccò M, Zhao Y, Williams P, Tian M, Tanalgo K, Woo K-S, Barjakovic M, McCracken GF, Simmons NB, Racey PA, Ford D, Labegalini JA, Colzato N, Pereira MJR, Aguiar LMS, Moratelli R, Preez GD, Pérez-González A, Reboleira ASPS, Gunn J, Cartney AM, Bobrowiec PED, Milko D, Kinuthia W, Fischer E, Meierhofer MB, & Frick WF. Brazilian cave heritage under siege. Science 375: 1238–1239. https://doi.org/10.1126/science.abo1973

113. Gladstone NS#, Niemiller ML, Hutchins B, Schwartz B, Czaja A, Slay ME, & Whelan NV. Subterranean freshwater gastropod biodiversity and conservation in the United States and Mexico. Conservation Biology 36: e13722. https://doi.org/10.1111/cobi.13722

112. Wynne JJ, Niemiller ML, & Chapin KJ. Evolutionary models influencing troglomorphy. Pp. 57–94 in: Cave Life – Drivers of Diversity and Diversification (Wynne JJ, ed.), John Hopkins Press.

111. Niemiller ML, Carter ET, Fenolio DB, Gluesenkamp AG, & Phillips JG. Drivers of subterranean colonization and diversification in cave-dwelling salamanders. Pp. 224–270 in: Cave Life – Drivers of Diversity and Diversification (Wynne JJ, ed.), John Hopkins Press.

110. Ober KA, Niemiller ML, & Philips TK. Cave trechine (Coleoptera: Carabidae) diversity and biogeography in North America. Pp. 192–223 in: Cave Life – Drivers of Diversity and Diversification (Wynne JJ, ed.), John Hopkins Press.

109. Zhao Y, Gluesenkamp AG, Fenolio DB, Soares D, Niemiller ML, Bichuette ME, & Chakrabarty P. Diversity, distribution, and conservation of cavefishes in China. Pp. 271–305 in: Cave Life – Drivers of Diversity and Diversification (Wynne JJ, ed.), John Hopkins Press.

2021

108. Niemiller ML, Carter ET, Gladstone NS#, Niemiller KDK, Hayter LE, Engel AS, Miller BT, & Fitzpatrick BM. The distribution, ecology, life history, and conservation status of the Berry Cave salamander (Gyrinophilus gulolineatus). Herpetological Conservation and Biology 16: 686–703.

107. Wynne JJ, Howarth FG, Mammola S, Ferreira RL, Cardoso P, Di Lorenzo T, Galassi DMP, Medellin RA, Miller BW, Sánchez-Fernández D, Bichuette ME, Biswas J, Boonyanusith C, Borges PAV, Boston PJ, Cheeptham N, Deharveng L, Eme D, Fenolio D, Fišer C, Fišer Z, Gon III S, Goudarzi F, Griebler C, Halse S, Hoch H, Katz AD, Kováč L, Lilley TM, Manchi S, Manenti R, Martínez A, Martinez WE, Meierhofer MB, Miller AZ, Moldovan OT, Niemiller ML, Pipan T, Pellegrini TG, Phillips-Lander CM, Poot C, Racey PA, de Rosário IA, Sendra A, Shear WA, Silva MS, Tian M, Taiti S, Venarsky MP, Pakarati SY, Zagmajster M, & Zhao Y. A roadmap for the conservation of the subterranean biome. Conservation Letters 14: e12834. https://doi.org/10.1111/conl.12834

106. Fitzgerald DB, Smith DR, Culver DC, Feller D, Fong DW, Hajenga J, Niemiller ML, Nolfi DC, Orndorff WD, Douglas B, Maloney KO, & Young JA. Using expert knowledge to support Endangered Species Act decision making for data-deficient species. Conservation Biology 35: 1627–1638. https://doi.org/10.1111/cobi.13694

105. Niemiller KDK, Davis MA, & Niemiller ML. Addressing biodiversity naivety using project-based learning with iNaturalist. Journal for Nature Conservation 64: 126070. https://doi.org/10.1016/j.jnc.2021.126070

104. Niemiller ML, Helf K, & Toomey RS. Mammoth Cave: a hotspot of subterranean biodiversity in the United States. Diversity 13: 373. https://doi.org/10.3390/d13080373

103. Mouser JB#, Brewer SK, Niemiller ML, Mollehauer R, & Van Den Bussche RA. Refining sampling protocols for cavefishes and cave crayfishes to account for environmental variation. Subterranean Biology 39: 79–105. https://doi.org/10.3897/subtbiol.39.64279

102. Gladstone NS#, Pieper EB, Keenan SW, Paterson AT, Slay ME, Dooley K#, Engel AS, & Niemiller ML. Discovery of the Blue Ridge springsnail, Fontigens orolibas, Hubricht, 1957 (Gastropoda: Emmericiidae) in east Tennessee and its conservation implications. Freshwater Mollusk Biology and Conservation 24: 34–42. https://doi.org/10.31931/fmbc-d-20-00007

101. Benito JB#, Porter ML, & Niemiller ML. The mitochondrial genomes of five spring and groundwater amphipods of the family Crangonyctidae (Crustacea: Amphipoda) from eastern North America. Mitochondrial DNA Part B 6: 1662–1667. https://doi.org/10.1080/23802359.2021.1926350

2020

100. Holler Jr C, Mays J, & Niemiller ML. The fauna of caves and other subterranean habitats of North Carolina, USA. Journal of Cave and Karst Studies 82: 221–260. https://doi.org/10.4311/2019LSC0133

99. Soares D, & Niemiller ML. Variation in cephalic neuromasts in surface and cave-dwelling fishes of the family Amblyopsidae (Teleostei: Percopsiformes). Journal of Cave and Karst Studies 82: 198–209. https://doi.org/10.4311/2019LSC0115

98. Boyd SH#, Niemiller KDK, Dooley KE#, Nix J^, & Niemiller ML. Using environmental DNA methods to survey for rare groundwater fauna: detection of an endangered endemic cave crayfish in northern Alabama. PLoS One 15: e0242741. https://doi.org/10.1371/journal.pone.0242741

97. Mammola S, Amorim IR, Bichuette ME, Borges P, Cheeptham N, Cooper SJB, Culver DC, Deharveng L, Eme D, Ferreira RL, Fiser C, Fiser Z, Fong DW, Griebler C, Jeffery WR, Kowalko JE, Jugovic J, Lilley TM, Malard F, Manenti R, Martinez A, Meierhofer MB, Northup DE, Pellegrini TG, Protas M, Niemiller ML, Reboleira AS, Pipan T, Venarsky MP, Wynne JJ, Zagmajster M, & Cardoso P. Fundamental research questions in subterranean biology. Biological Reviews 95: 1855–1872. https://doi.org/10.1111/brv.12642

96. Zigler KS, Niemiller ML, Stephen CDR#, Ayala BN, Milne MA, Gladstone NS#, Engel AS, Jensen JB, & Ozier J. Cave biodiversity of Georgia. Journal of Cave and Karst Studies 82: 125–167. https://doi.org/10.4311/2019LSC0125

95. Baker SJ, Niemiller ML, Stites AJ, Ash KT, Davis MA, Dreslik MJ, & Phillips CA. Evaluation of environmental DNA to detect Sistrurus catenatus and Ophidiomyces ophiodiicola in crayfish burrows. Conservation Genetics Resources 12: 13–15. https://doi.org/10.1007/s12686-018-1053-9

94. DiStefano RJ, Ashley D, Brewer SK, Mouser JB#, & Niemiller ML. Preliminary investigation of the critically imperiled Caney Mountain Cave Crayfish Orconectes stygocaneyi (Decapoda: Cambaridae) (Hobbs III 2001) in Missouri, USA. Freshwater Crayfish 25: 47–57. http://dx.doi.org/10.5869/fc.2020.v25-1.047

93. Hart PB#, Niemiller ML, Burress ED, Armbruster JW, Ludt WB, & Chakrabarty P. Cave-adapted evolution in the North American amblyopsid fishes inferred using phylogenomics and geometric morphometrics. Evolution 74: 936–949. https://doi.org/10.1111/evo.13958

92. Soares D, & Niemiller ML. Extreme adaptation in caves. The Anatomical Record 303: 15–23. https://doi.org/10.1002/ar.24044

2019

91. Gladstone NS#, Perez KE, Pieper EB^, Carter ET, Dooley KE#, Shoobs NF#, Engel AS, & Niemiller ML. A new species of stygobitic snail in the genus Antrorbis Hershler and Thompson, 1990 (Gastropoda, Cochliopidae) from the Appalachian Valley and Ridge of eastern Tennessee, USA. ZooKeys 898: 103–120. https://doi.org/10.3897/zookeys.898.46917

90. Niemiller ML, Inebnit T, Hinkle A, Jones B, Jones M, Lamb J#, Mann N, Miller B, Pinkley J, Pitts S, Sapkota K, & Slay ME. Discovery of a new population of the federally endangered Alabama cave shrimp, Palaemonias alabamae Smalley, 1961, in northern Alabama. Subterranean Biology 32: 43–59. https://doi.org/10.3897/subtbiol.32.38280

89. Wollenberg Valero KC, Marshall JC, Bastiaans E, Caccone A, Camargo A, Morando M, Niemiller ML, Pabijan M, Russello MA, Sinervo B, Werneck JFP, Sites Jr JW, Wiens JJ, & Steinfartz S. Patterns, mechanisms, and genetics of speciation in reptiles and amphibians. Genes 10: 646. https://doi.org/10.3390/genes10090646

88. Mammola S, Cardoso P, Culver DC, Deharveng L, Ferreira RL, Fiser C, Galassi DMP, Griebler C, Halse S, Humphreys WF, Isaia M, Malard F, Martinez A, Moldovan OT, Niemiller ML, Pavlek M, Reboleira ASPS, Souza-Silva M, Teeling EC, Wynne JJ, & Zagmajster M. Scientists’ warning on the conservation of subterranean ecosystems. BioScience 69: 641–650. https://doi.org/10.1093/biosci/biz064

87. Gladstone NS#, Niemiller ML, Pieper EB^, Dooley KE^, & McKinney ML. Morphometrics and phylogeography of the cave-obligate land snail Helicodiscus barri (Gastropoda, Stylommatophora, Helicodiscidae). Subterranean Biology 30: 1–32. https://doi.org/10.3897/subtbiol.30.35321

86. Niemiller ML, & Taylor SJ. Protecting cave life. Pp. 822–829 in: Encyclopedia of Caves, 3rd edition (Culver DC, White WB, & Pipan T, eds.). Elsevier. https://doi.org/10.1016/B978-0-12-814124-3.00099-6

85. Niemiller ML, Taylor SJ, Slay ME, & Hobbs III HH. Biodiversity in the United States and Canada. Pp. 163–176 in: Encyclopedia of Caves, 3rd edition (Culver DC, White WB, & Pipan T, eds.). Elsevier. https://doi.org/10.1016/B978-0-12-814124-3.00018-2

84. Niemiller ML, Bichuette ME, Chakrabarty P, Fenolio DB, Gluesenkamp AG, Soares D, & Zhao Y. Cavefishes. Pp. 227–236 in: Encyclopedia of Caves, 3rd edition (Culver DC, White WB, & Pipan T, eds.). Elsevier. https://doi.org/10.1016/B978-0-12-814124-3.00026-1

83. Goricki S, Niemiller ML, Fenolio DB, & Gluesenkamp AG. Salamanders. Pp. 871–884 in: Encyclopedia of Caves, 3rd edition (Culver DC, White WB, & Pipan T, eds.). Elsevier. https://doi.org/10.1016/B978-0-12-814124-3.00104-7

2018

82. Gladstone NS#, Carter ET, Niemiller KDK, Hayter LE, & Niemiller ML. A new maximum body size record for the Berry Cave salamander (Gyrinophilus gulolineatus) and genus Gyrinophilus (Caudata, Plethodontidae) with a comment on body size in plethodontid salamanders. Subterranean Biology 28: 29–38. https://doi.org/10.3897/subtbiol.28.30506

81. Gladstone NS#, Carter ET, McKinney ML, & Niemiller ML. Status and distribution of cave-obligate land snails in the Appalachians and Interior Low Plateau of the eastern United States. American Malacological Bulletin 36: 62–78. https://doi.org/10.4003/006.036.0107

80. Niemiller ML, Porter ML, Keany J#, Gilbert H, Fong D, Culver DC, Kendall KD, Davis MA, & Taylor SJ. Evaluation of eDNA for groundwater invertebrate detection and monitoring: a case study with endangered Stygobromus (Amphipoda: Crangonyctidae). Conservation Genetics Resources 10: 247–257. https://doi.org/10.1007/s12686-017-0785-2

79. Niemiller ML, Bichuette E, & Taylor SJ. Conservation of cave fauna in Europe and the Americas. Pp. 451–478 in: Ecological Studies: Cave Ecology (Moldovan OT, Kovac L, & Halse S, eds). Springer. http://dx.doi.org/10.1007/978-3-319-98852-8_22

78. Niemiller ML, Slay ME, & Bitting CJ, eds. Proceedings of the 22nd National Cave and Karst Management Symposium, 16–20 October 2017, Eureka Springs, Arkansas. 106 pp.

2017

77. Soares D, Adams R#, Hammond S, Slay ME, Fenolio DB, & Niemiller ML. Evolution of coprophagy and nutrient absorption in a cave salamander. Subterranean Biology 24: 1–9. https://doi.org/10.3897/subtbiol.24.15013

76. Nelson SK, Niemiller ML, & Fitzpatrick BM. Co-occurrence and hybridization between Necturus maculosus and an undescribed Necturus species in the Southern Appalachians. Journal of Herpetology 51: 559–566. https://doi.org/10.1670/17-006

75. Niemiller ML, Zigler KS, Ober KA, Carter ET#, Engel AS, Moni G, Philips TK, & Stephen CDR#. Rediscovery and conservation status of six short-range endemic Pseudanophthalmus cave beetles (Carabidae: Trechini). Insect Conservation and Diversity 10: 495–501. https://doi.org/10.1111/icad.12263

74. Fenolio DB, Niemiller ML, Gluesenkamp AG, McKee AM, & Taylor SJ. New distributional records of the stygobiotic crayfish Cambarus cryptodytes (Decapoda: Cambaridae) in the Floridan aquifer system of southwestern Georgia. Southeastern Naturalist 16: 163–181. https://doi.org/10.1656/058.016.0205

73. Bagley R#, Sousa V, Niemiller ML, & Linnen C. History, geography, and host use shape genome-wide patterns of genetic variation in the redheaded pine sawfly (Neodiprion lecontei). Molecular Ecology 26: 1022–1044. https://doi.org/10.1111/mec.13972

72. Engel AE, Paterson AT, & Niemiller ML. Biogeography of microbes from caves in the Interior Low Plateau and Appalachian karst regions. Pp. 410 in: Proceedings of the 17th International Congress of Speleology, Volume 1 (Moore K, & White S). Australian Speleological Federation & Speleo2017.

71. Keenan SW, Paterson AT, Niemiller ML, Slay ME, Clark SA, & Engel AS. Observations of the first stygobiont snail (Hydrobiidae, Fontigens sp.) in Tennessee. Pp. 91–94 in: Proceedings of the 17th International Congress of Speleology, Volume 1 (Moore K, & White S). Australian Speleological Federation & Speleo2017.

70. Engel AS, Niemiller ML, Zigler KS, Stephen CDR, Carter ET, Paterson AT, Keenan SW, & Taylor SJ. Invertebrate and vertebrate cave fauna records for the Appalachian Valley and Ridge. Pp. 82–86 in: Proceedings of the 17th International Congress of Speleology, Volume 1 (Moore K, & White S). Australian Speleological Federation & Speleo2017.

2016

69. Niemiller ML, Zigler KS, Hart PB#, Kuhajda BR, Ayala BN^, Armbruster JW, & Engel AS. First record of a stygobiotic fish (Percopsiformes: Amblyopsidae: Typhlichthys) from the Appalachians karst region in the eastern United States. Subterranean Biology 20: 39–50. https://doi.org/10.3897/subtbiol.20.9693

68. Armbruster JW, Niemiller ML, & Hart PB#. Morphological evolution of the cave-, spring- and swampfishes of the family Amblyopsidae (Percopsiformes). Copeia 104: 763–777. https://doi.org/10.1643/CI-15-339

67. Christman MC, Doctor DH, Niemiller ML, Weary DJ, Young JA, Zigler KS, & Culver DC. Predicting the occurrence of cave-inhabiting fauna based on features of the Earth surface environment. PLoS One 11: e0160408. https://doi.org/10.1371/journal.pone.0160408

66. Reynolds RG, Collar DC, Pasachnik SA, Niemiller ML, Puente-Rolon AR, & Revell LJ. Ecological specialization and morphological diversification in Greater Antillean boas. Evolution 70: 1882–1895. https://doi.org/10.1111/evo.12987

65. Niemiller ML, Zigler KS, Stephen CDR#, Carter ET#, Paterson AT, Taylor SJ, & Engel AS. Vertebrate fauna in caves of eastern Tennessee within the Appalachians karst region, USA. Journal of Cave and Karst Studies 78: 1–24. https://doi.org/10.4311/2015lsc0109

64. Niemiller ML, Glorioso BM, Fenolio DB, Reynolds RG, Taylor SJ, & Miller BT. Growth, survival, longevity, and population size of the Big Mouth Cave salamander (Gyrinophilus palleucus necturoides) from the type locality in Grundy Co., Tennessee, USA. Copeia 104: 35–41. https://doi.org/10.1643/OT-14-197

63. Taylor SJ, & Niemiller ML. Biogeography and conservation of Bactrurus groundwater amphipods (Crangonyctidae) in the central and eastern United States. Subterranean Biology 17: 1–29. https://doi.org/10.3897/subtbiol.17.7298

62. Soares D, Niemiller ML, & Higgs DM. Hearing in cavefishes. Pp. 187–195 in: Fish Hearing and Bioacoustics – An Anthology in Honor of Arthur N. Popper and Richard R. Fay (Sisneros JA, ed). Springer. https://doi.org/10.1007/978-3-319-21059-9_9

61. Slay ME, Niemiller ML, Taylor SJ, & Sutton M. Cave Life of the Ozarks: A Guide to Commonly Encountered Species in Arkansas, Missouri, and Oklahoma. Biology Section of the National Speleological Society, Huntsville, Alabama. 45 pp.

2015

60. Taylor SJ, Krejca JK, Niemiller ML, Dreslik MJ, & Phillips CA. Life history and demographic differences between cave and surface populations of the western slimy salamander Plethodon albagula (Caudata: Plethodontidae) in central Texas. Herpetological Conservation and Biology 10: 740–752.

59. Niemiller ML*, & Soares D*. Cave environments. Pp. 161–191 in: Extremophile Fishes – Ecology and Evolution of Teleosts in Extreme Environments (Riesch R, Plath M, & Tobler M, eds). Springer. https://doi.org/10.1007/978-3-319-13362-1_8

2014

58. Thompson S, Muzinic L, Niemiller ML, & Voss SR. Probability of regenerating a normal limb after bite injury in the Mexican axolotl (Ambystoma mexicanum). Regeneration 1: 27–32. https://doi.org/10.1002/reg2.17

57. Kendall KD, Niemiller ML, Dittrich-Reed D, & Schussler EE. Helping graduate teaching assistants in biology use student evaluations as professional development. The American Biology Teacher 76: 584–588. https://doi.org/10.1525/abt.2014.76.9.3

56. Fenolio DB, Niemiller ML, Bonett RM, Graening GO, Collier BA, & Stout JF. Life history, demography, and the influence of cave-roosting bats on a population of the grotto salamander (Eurycea spelaea) from the Ozark Plateaus of Oklahoma (Caudata: Plethodontidae). Herpetological Conservation and Biology 9: 394–405.

55. Espinasa L, Espinasa M, Fenolio DB, Slay ME, & Niemiller ML. Distribution and conservation status of Speleonycta ozarkensis (Insecta: Zygentoma: Nicoletiidae) from caves of the Ozark Highlands of Arkansas and Oklahoma, USA. Subterranean Biology 14: 51–62. https://doi.org/10.3897/subtbiol.14.8275

54. Chakrabarty P, Prejean JA, & Niemiller ML. The Hoosier cavefish, a new and endangered species (Amblyopsidae, Amblyopsis) from the caves of southern Indiana. ZooKeys 412: 41–57. https://doi.org/10.3897/zookeys.412.7245

53. Soares D, Niemiller ML, & Higgs D. Hearing and acoustic communication in cavefishes. Ambient Science 1: 1–6.

52. Kennan SW, Niemiller ML, & Williams BW. Observations of Cambarus bartonii cavatus (Decapoda: Cambaridae) and ectosymbiotic branchiobdellidans (Annelida: Clitellata) in Cruze Cave, Knox County, Tennessee, USA. Speleobiology Notes 6: 55–61.

51. Fenolio DB, Niemiller ML, & Martinez B. Observations of reproduction in captivity by the Dougherty Plain Cave Crayfish, Cambarus cryptodytes (Decapoda: Astacoidea: Cambaridae). Speleobiology Notes 6: 14–26.

50. Reynolds RG, Niemiller ML, & Revell LJ. Toward a Tree-of-Life for the boas and pythons: multilocus species-level phylogeny with unprecedented taxon sampling. Molecular Phylogenetics and Evolution 71: 201–213. https://doi.org/10.1016/j.ympev.2013.11.011

49. Niemiller ML, & Reeves WC. Predation by the stygophilic crayfish Cambarus tenebrosus on the salamander Pseudotriton ruber within a cave in Cannon County, Tennessee, USA. Speleobiology Notes 6: 8–13.

2013

48. Barun A, Niemiller ML, Fitzpatrick BM, Fordyce JA, & Simberloff D. Can genetic data confirm or refute historical records: the island invasion of the small Indian mongoose (Herpestes auropunctatus). Biological Invasions 15: 2243–2251. https://doi.org/10.1007/s10530-013-0447-6

47. Fenolio DB, Niemiller ML, Levy M, & Martinez B. Conservation status of the Georgia blind salamander (Eurycea wallacei) from the Floridan Aquifer of Florida and Georgia. Reptiles and Amphibians 20: 97–111. https://doi.org/10.17161/randa.v20i3.13945      

46. Kendall KD, Niemiller ML, Dittrich-Reed D, Chick LD, Wilmoth L, Milt A, Burt M, Lopes N, Cantwell L, Rubio L, Allison A, & Schussler EE. Departments can develop teaching identities of graduate students. CBE – Life Sciences Education 12: 316–317. https://doi.org/10.1187/cbe.13-03-0066

45. McAllister CT, Bursey CR, Fenolio DB, & Niemiller ML. Bothriocephalus sp. (Cestoidea: Bothriocephalidea: Bothriocephalidae) from the Georgia blind salamander, Eurycea wallacei (Caudata: Plethodontidae), in Georgia, USA: first definitive report of a parasite from this host. Comparative Parasitology 80: 308–311. https://doi.org/10.1654/4615.1

44. Niemiller ML, Graening GO, Fenolio DB, Godwin JC, Cooley JR, Pearson WR, Near TJ, & Fitzpatrick BM. Doomed before they are described? The need for conservation assessments of cryptic species complexes using an amblyopsid cavefish (Amblyopsidae: Typhlichthys) as a case study. Biodiversity and Conservation 22: 1799–1820. https://doi.org/10.1007/s10531-013-0514-4

43. Reynolds RG, Niemiller ML, Hedges SB, Dornburg A, & Revell LJ. Molecular phylogeny and historical biogeography of West Indian boid snakes (Chilabothrus). Molecular Phylogenetics and Evolution 68: 461–470. https://doi.org/10.1016/j.ympev.2013.04.029

42. Niemiller ML, & Zigler KS. Patterns of cave biodiversity and endemism in the Appalachians and Interior Plateau of Tennessee USA. PLoS One 8: e64177. https://doi.org/10.1371/journal.pone.0064177

41. Fenolio DB, Niemiller ML, Soares D, Slay ME, Harris KA, & Harris N. Subterranean reproduction of the ringed crayfish, Orconectes neglectus Faxon 1885 (Astacoidea: Cambaridae) within a cave in the Ozark Highlands of Oklahoma, USA. Speleobiology Notes 5: 43–46.

40. Fenolio DB, McAllister CT, Niemiller ML, Soares D, & Cooley J. An extreme case of trematode parasitic infection of a larval Ozark blind cave salamander, Eurycea spelaea (Caudata: Plethodontidae) from the Ozark Highlands of Missouri, USA. Speleobiology Notes 5: 34–37.

39. Fenolio DB, Zhao Y, Niemiller ML, & Stout JF. In situ observations of seven enigmatic cave loaches and one cave barbel from Guangxi, China, with notes on conservation status. Speleobiology Notes 5: 19–33.

38. Fenolio DB, Niemiller ML, Soares D, Slay ME, Stark RC, & Hensley SL. A maximum size and maximum abundance record of the Delaware County cave crayfish, Cambarus subterraneus Hobbs 1993 (Astacoidea: Cambaridae), from Twin Cave, Delaware County, Oklahoma, USA. Speleobiology Notes 5: 9–13.

37. Soares D, & Niemiller ML. Sensory adaptations of cavefishes to cave environments. BioScience 63: 274–283. https://doi.org/10.1525/bio.2013.63.4.7

36. Niemiller ML, McCandless JR, Reynolds RG, Caddle J, Tillquist CR, Near TJ, Pearson WD, & Fitzpatrick BM. Effects of climatic and geological processes during the Pleistocene on the evolutionary history of the northern cavefish, Amblyopsis spelaea (Teleostei: Amblyopsidae). Evolution 67: 1011–1025. https://doi.org/10.1111/evo.12017

35. Niemiller ML, Higgs DM, & Soares D. Evidence for hearing loss in amblyopsid cavefishes. Biology Letters 9: 20130104. https://doi.org/10.1098/rsbl.2013.0104

34. Niemiller ML, Fitzpatrick BM, Shah P, Schmitz L, & Near TJ. Evidence for repeated loss of selective constraint in rhodopsin of amblyopsid cavefishes. Evolution 67: 732–748. https://doi.org/10.1111/j.1558-5646.2012.01822.x

33. Niemiller ML, Zigler KS, & Fenolio DB. Cave Life of TAG: A Guide to Commonly Encountered Species in Tennessee, Alabama and Georgia. Biology Section of the National Speleological Society, Huntsville, Alabama. 45 pp.

32. Niemiller ML, Reynolds RG, & Miller BT (eds.). The Reptiles of Tennessee. University of Tennessee Press, Knoxville, Tennessee. 347 pp.

2012

31. Reynolds RG, Niemiller ML, & Fitzpatrick BM. Genetic analysis of an endemic archipelagic lizard reveals sympatric cryptic lineages and taxonomic discordance. Conservation Genetics 13: 953–963. https://doi.org/10.1007/s10592-012-0344-z

30. Miller BT, & Niemiller ML. Gyrinophilus palleucus. Catalogue of American Amphibians and Reptiles 884: 1–7.

29. Niemiller ML, Near TJ, & Fitzpatrick BM. Delimiting species using multilocus data: diagnosing cryptic diversity in the southern cavefish Typhlichthys subterraneus (Teleostei: Amblyopsidae). Evolution 66: 846–866. https://doi.org/10.1111/j.1558-5646.2011.01480.x

28. Fitzpatrick BM, Fordyce JA, Niemiller ML, & Reynolds RG. What can DNA tell us about biological invasions? Biological Invasions 14: 245–253. https://doi.org/10.1007/s10530-011-0064-1

27. Goricki A, Niemiller ML, & Fenolio DB. Salamanders. Pp. 665–676 in: Encyclopedia of Caves, 2nd Edition (White WH, & Culver DC, eds.). Elsevier. https://doi.org/10.1016/b978-0-12-383832-2.00098-0

2011

26. Niemiller ML, Reynolds RG, Glorioso BM, Spiess J, & Miller BT. Herpetofauna of the cedar glades and associated habitats of the Inner Central Basin of middle Tennessee. Herpetological Conservation and Biology 6: 135–149.

25. Reynolds RG, Niemiller ML, & Pasachnik SA. Distribution of the Pygmy Salamander (Desmognathus wrighti) in Tennessee. Journal of the Tennessee Academy of Science 86: 53–55.

24. Niemiller ML, & Reynolds RG (eds.). The Amphibians of Tennessee. University of Tennessee Press, Knoxville, Tennessee. 369 pp.

2010

23. Glorioso BM, Niemiller ML, & Cobb VA. Feeding activity of Amphiuma tridactylum at Reelfoot Lake, Tennessee. Journal of the Tennessee Academy of Science 85: 87–90.

22. Reynolds RG, & Niemiller ML. 2010. Island invaders: introduced reptiles and amphibians of the Turks and Caicos Islands. Reptiles and Amphibians 17: 117–121.

21. Niemiller ML, Osbourn MS, Fenolio DB, Pauley TK, Miller BT, & Holsinger JR. Conservation status and habitat use of the West Virginia Spring Salamander (Gyrinophilus subterraneus) and Spring Salamander (G. porphyriticus) in General Davis Cave, Greenbrier Co., West Virginia. Herpetological Conservation and Biology 5: 32–43.

20. Niemiller ML, & Miller BT. Gyrinophilus gulolineatus. Catalogue of American Amphibians and Reptiles. 862.1.

19. Niemiller ML, Nosil P, & Fitzpatrick BM. Corrigendum. Recent divergence-with-gene-flow in Tennessee cave salamanders (Plethodontidae: Gyrinophilus) inferred from gene genealogies. Molecular Ecology 19: 1513–1514. https://doi.org/10.1111/j.1365-294x.2010.04564.x

18. Graening GO, Fenolio DB, Niemiller ML, Brown AV, & Beard JB. The 30-year recovery effort for the Ozark cavefish (Amblyopsis rosae): analysis of current distribution, population trends, and conservation status of this threatened species. Environmental Biology of Fishes 87: 55–88. https://doi.org/10.1007/s10641-009-9568-2

17. Niemiller ML, & Poulson TL. Subterranean fishes of North America: Amblyopsidae. Pp. 169–280 in: The Biology of Subterranean Fishes (Trajano E, Bichuette ME, & Kapoor BG, eds). Science Publishers, Enfield, New Hampshire, USA. https://doi.org/10.1201/ebk1578086702

2009

16. Niemiller ML, Glorioso BM, Nicholas C, Phillips J, Rader J, Reed E, Sykes KL, Todd J, Wyckoff GR, Young EL, & Miller BT. Notes on the reproduction of the streamside salamander, Ambystoma barbouri, from Rutherford County, Tennessee. Southeastern Naturalist 8: 37–44. https://doi.org/10.1656/058.008.0104

15. Niemiller ML, Fenolio D, Graening GO, & Miller BT. Observations on oviposition and reproduction of the cave salamander, Eurycea lucifuga (Caudata: Plethodontidae), from Arkansas and Tennessee. Speleobiology Notes 1: 17–19.

14. Niemiller ML, Miller BT, & Fitzpatrick BM. Systematics and evolutionary history of subterranean Gyrinophilus salamanders. Proceedings of the 15th International Congress of Speleology, Kerrville, Texas 15: 242–248.

13. Niemiller ML, & Miller BT. A survey of the cave-associated amphibians of the eastern United States with an emphasis on salamanders. Proceedings of the 15th International Congress of Speleology, Kerrville, Texas 15: 249–256.

2008

12. Miller BT, Niemiller ML, & Reynolds RG. Observations on egg-laying behavior and interactions among attend female red salamanders (Pseudotriton ruber) with comments on the use of caves by this species. Herpetological Conservation and Biology 3: 203–210.

11. Fitzpatrick BM, Placyk Jr JS, Niemiller ML, Casper GS, & Burghardt GM. Distinctiveness in the face of gene flow: hybridization between specialist and generalist gartersnakes. Molecular Ecology 17: 4107–4117. https://doi.org/10.1111/j.1365-294x.2008.03885.x

10. Niemiller ML, Fitzpatrick BM, & Miller BT. Recent divergence-with-gene-flow in Tennessee cave salamanders (Plethodontidae: Gyrinophilus) inferred from gene genealogies. Molecular Ecology 17: 2258–2275. https://doi.org/10.1111/j.1365-294x.2008.03750.x

9. Miller BT, & Niemiller ML. Distribution and relative abundance of Tennessee cave salamanders (Gyrinophilus palleucus and G. gulolineatus) with an emphasis on Tennessee populations. Herpetological Conservation and Biology 3: 1–20.

8. Niemiller ML, & Fitzpatrick BM. Phylogenetics of the southern cavefish, Typhlichthys subterraneus: implications for conservation and management. Proceedings of the 18th National Cave and Karst Management Symposium 18: 79–88.

2007

7. McAllister CT, Bursey CR, Niemiller ML, & Miller BT. A noteworthy infection of Clinostomum complanatum (Digenea: Clinostomidae) in a cave salamander, Eurycea lucifuga (Caudata: Plethodontidae), from northcentral Tennessee. Texas Journal of Science 59: 321–326.

6. Niemiller ML, & Miller BT. Subterranean reproduction of the southern two-lined salamander (Eurycea cirrigera) from Short Mountain, Tennessee. Herpetological Conservation and Biology 2: 106–112.

2006

5. Niemiller ML, Glorioso BM, Nicholas C, Phillips J, Rader J, Reed E, Sykes KL, Todd J, Wyckoff GR, Young EL, & Miller BT. Status and distribution of the streamside salamander (Ambystoma barbouri) in middle Tennessee. American Midland Naturalist 156: 393–399. https://doi.org/10.1674/0003-0031(2006)156[394:sadots]2.0.co;2

4. Glorioso BM, & Niemiller ML. Using deep-water crawfish nets to capture aquatic turtles. Herpetological Review 37: 185–187.

3. Niemiller ML, Glorioso BM, & Miller BT. An eastern and subterranean maximum size record of the banded sculpin, Cottus carolinae, from Tennessee. Journal of the Tennessee Academy of Science 81: 41–43.

2005

2. Koczaja C, McCall L, Fitch E, Glorioso B, Hanna C, Kyzar J, Niemiller M, Spiess J, Tolley A, Wyckoff R, & Mullen D. Size-specific habitat segregation and intraspecific interactions in banded sculpin (Cottus carolinae). Southeastern Naturalist 4: 207–218. https://doi.org/10.1656/1528-7092(2005)004[0207:shsaii]2.0.co;2

1. Niemiller ML. The herpetofauna of the upper Duck River watershed in Coffee County, Tennessee. Journal of the Tennessee Academy of Science 80: 6–12.


Short Notes

2024

33. Quering CI^, Thomas WR^, Bell KE#, Bwalya PK^, Driver PN^, Jones AP^, Lee HK^, Long BA^, Richards ML#, Niemiller GA, Niemiller ER, Niemiller KDK, & Niemiller ML. Geographic distribution. Hemidactylium scutatum. Herpetological Review 55: 214.

2022

32. Niemiller ML, Niemiller ER, & Niemiller KDK. Geographic distribution. Farancia abacura. Herpetological Review 53: 445–446.

2021

31. Niemiller ML, Niemiller ER, & Niemiller KDK. Geographic distribution. Nerodia rhombifer. Herpetological Review 52: 579.

30. Gaulke SM, Chace JZ, Taylor SJ, Davis MA, Niemiller ML, & Schuett GW. Agkistrodon contortrix (Eastern Copperhead). Reproduction, male combat, and phenology. Herpetological Review 52: 414–415.

29. Swain MT & Niemiller ML. Gyrinophilus porphyriticus (Spring Salamander). Diet. Herpetological Review 52: 108–109.

2020

28. Gladstone NS#, Royalty TM^, Carter ET, & Niemiller ML. Eurycea lucifuga (Cave Salamander). Diet. Herpetological Review 51: 291.

2012

27. Nelson SK^, Niemiller ML, & Reynolds RG. New county records for amphibians and reptiles in Tennessee. Herpetological Review 43: 456–457.

2011

26. Reynolds, RG, & Niemiller ML. Epicrates chrysogaster chrysogaster (Turks Island Boa). Diet. Herpetological Review 42: 290.

25. Reynolds RG, Niemiller ML, & Manco BN. Epicrates chrysogaster chrysogaster (Turks Island Boa). Record Size. Herpetological Review 42: 290.

24. Reynolds RG, Niemiller ML, Riggs BM, & Manco BN. Geographic distribution. Osteopilus septentrionalis. Herpetological Review 42: 237–238.

2010

23. Reynolds RG, & Niemiller ML. Distribution. Epicrates chrysogaster (Southern Bahamas Boa). Caribbean Herpetology 1: 14.

2009

22. Reynolds RG, & Niemiller ML. Geographic distribution. Hemidactylus mabouia. Herpetological Review 40: 452.

21. Niemiller ML, Reynolds RG, Reynolds JG, & Miller BT. Gyrinophilus porphyriticus porphyriticus. Reproduction. Herpetological Review 40: 67.

2008

20. Niemiller ML, & Miller BT. Plethodon dorsalis. Reproduction. Herpetological Review 39: 335.

2007

19. Niemiller ML, Glorioso BM, & Wyckoff GR. New county records for amphibians in middle Tennessee. Herpetological Review 38: 234.

18. Niemiller ML, Glorioso BM, Young EL, & Cobb VA. Farancia abacura reinwardtii. Reproduction. Herpetological Review 38: 207.

2006

17. Niemiller ML, Miller JA, Miller JH, & Miller BT. Geographic distribution. Lampropeltis calligaster. Herpetological Review 37: 498.

16. Miller JH, Miller JA, Niemiller ML, & Miller BT. Geographic distribution. Rana palustris. Herpetological Review 37: 490.

15. Niemiller ML, & Miller BT. Geographic distribution. Rana palustris. Herpetological Review 37: 489–490.

14. Miller JA, Miller JH, Niemiller ML, & Miller BT. Geographic distribution. Bufo fowleri. Herpetological Review 37: 486.

13. Glorioso BM & Niemiller ML. Geographic distribution. Plethodon glutinosus. Herpetological Review 37: 485.

12. Niemiller ML, & Miller BT. Geographic distribution. Plethodon dorsalis. Herpetological Review 37: 484.

11. Niemiller ML, & Miller BT. Geographic distribution. Eurycea lucifuga. Herpetological Review 37: 483.

10. Niemiller ML, and Miller BT. Geographic distribution. Eurycea cirrigera. Herpetological Review 37: 483.

9. Niemiller ML, Glorioso BM, Gray E, Miller BT, Jensen JB, & Keyes T. Pseudotriton ruber ruber. Size and subterranean aggregation. Herpetological Review 37: 438.

8. Niemiller ML, & Glorioso BM. Geographic distribution. Rana palustris. Herpetological Review 37: 102.

7. Niemiller ML, Todd JM, Miller BT, & Mann NG. Geographic distribution. Gastrophryne carolinensis. Herpetological Review 37: 100.

6. Glorioso BM, & Niemiller ML. Geographic distribution. Bufo americanus americanus. Herpetological Review 37: 98.

5. Glorioso BM, Young EL, & Niemiller ML. Geographic distribution. Ambystoma talpoideum. Herpetological Review 37: 97.

2005

4. Niemiller ML, & Miller BT. Rana clamitans melanota. Predation. Herpetological Review 36: 440.

3. Miller BT, & Niemiller ML. Psuedotriton ruber ruber. Reproduction. Herpetological Review 36: 429.

2004

1. Niemiller ML. Geographic distribution. Gyrinophilus porphyriticus. Herpetological Review 35: 76.


Technical Reports

2024

39. Niemiller ML, Maxwell E#, & Higgs JP#. A biological inventory of Sinking Spring Cave at Abraham Lincoln Birthplace National Historical Park, Larue County, Kentucky, USA. Technical report no. 2024-04 to Western Kentucky University & Abraham Lincoln Birthplace National Historical Park, National Park Service, Department of Interior. 21 pp.

38. Niemiller ML, Higgs J#, Peterson B#, & Niemiller KDK. Pathogen screening of the Tennessee Cave Salamander (Gyrinophilus palleucus) to guide conservation and management in Tennessee. Technical report no. 2024-03 to Tennessee Wildlife Resources Agency. 19 pp.

37. Niemiller ML, Hinkle A, Cramphorn B#, Higgs J#, Maxwell E#, Peterson B#, Zigler KS, & Niemiller KDK. Assessing the distribution, pathogen screening, and status of the Tennessee Cave Salamander (Gyrinophilus palleucus) to guide conservation and management. Technical report no. 2024-02 to Tennessee Wildlife Resources Agency. 59 pp.

36. Niemiller ML, Higgs JP#, Peterson B#, & Niemiller KDK. Pathogen screening of the Tennessee Cave Salamander (Gyrinophilus palleucus) to guide conservation and management in Alabama. Technical report no. 2024-01 to Alabama Department of Conservation and Natural Resources. 19 pp.

35. Tobin B, Inebnit T, Niemiller ML, Slay M, & Miller B. Comprehensive biodiversity inventory and hydrological analysis of the Key Cave System at Key Cave National Wildlife Refuge. Report to U.S. Fish and Wildlife Service, Wheeler National Wildlife Refuge, Decatur, Alabama. 31 pp.

2023

34. Cucalón RV#, Corush JB, Niemiller ML, Davis MA, & Tan M. Species distributions and population structure of Illinois’ spring cavefish (Forbesichthys spp.) using genomics. Technical report to Illinois Department of Natural Resources. 27 pp.

33. Niemiller ML, & Higgs JP#. Environmental DNA analysis of groundwater in Benton County, Arkansas for the Ozark Cavefish (Troglichthys rosae). Technical report no. 2023-02 to ECO, Inc. 8 pp.

32. Niemiller ML, Hinkle A, Cramphorn B#, Higgs J#, Maxwell E#, Peterson B#, & Niemiller KDK. Assessing the distribution, pathogen screening, and status of the Tennessee Cave Salamander (Gyrinophilus palleucus) to guide conservation and management. Technical report no. 2023-01 to Alabama Department of Conservation and Natural Resources. 60 pp.

2022

31. Miller BV, & Niemiller ML. Seepage investigations, cave mapping, and a cave bioinventory conducted in Wear Cove, Tennessee. Restricted-File Federal Interagency Report prepared for use by the National Park Service. 63 pp.

2021

30. Niemiller ML, Niemiller KDK, Dooley KE#, Giltner K^, & Guillemette A^. Using environmental DNA to detect and monitor Alabama Cavefish and Alabama Cave Shrimp. Alabama Department of Conservation & Natural Resources, Montgomery, Alabama. 28 pp.

2020

29. Niemiller ML, & Niemiller KDK. Species status assessment for the Tennessee Cave Salamander (Gyrinophilus palleucus) McCrady, 1954. Version 1.0. Tennessee Wildlife Resources Agency, Nashville, Tennessee. 59 pp.

28. Davis MA, Molano-Flores B, Niemiller ML, Harper LR, & Benito J#. Assessing pollinator communities via an environmental DNA metacommunity assay. SERDP RC19-1102. Department of Defense, Strategic Environmental Research and Development Program. 59 pp.

27. Niemiller ML, Hardman R, Thames D, Istvanko D, Ogle C, Davis MA, Tan M, Apodaca JJ, Benito J#, Campbell J, Clark T, Dooley K#, Miller D, Niemiller KDK, Simpson C, & Upchurch P. Assessment of geographic range, habitat, and population genetics of the green salamander (Aneides aeneus) in Tennessee. Southeastern Association of Fish & Wildlife Agencies, South Carolina. 86 pp.

2018

26. Niemiller ML, Carter ET, Gladstone NS#, Hayter LE, & Angel AS. New surveys and reassessment of the conservation status of the Berry Cave Salamander (Gyrinophilus gulolineatus). United States Fish & Wildlife Service, Cookeville, Tennessee. 58 pp.

25. Casper, G.S., & Niemiller, M.L. 2018. Chapter 3.3 Bat Community Assessment. Pages 1–43: 4 appendices. In Casper, G.S. & Robson, J.L. Milwaukee Estuary Area of Concern Wildlife Population Assessment Report. Technical report to Wisconsin Department of Natural Resources, Office of Great Waters.

24. Hohoff TC, Niemiller ML, & Dreslik MJ. 2017 bat monitoring along the I-90 corridor. Illinois Natural History Survey Technical Report 2018(4): 1–8.

2017

23. Niemiller ML, Hohoff T, & Dreslik MJ. Habitat assessment for northern long-eared bats (Myotis septentrionalis) and Indiana bats (Myotis sodalis) at I-88 in Lee County, Illinois. Illinois Natural History Survey Memorandum ME-17-01. 20 pp.

22. Niemiller ML, & Dreslik MJ. Post-construction acoustic monitoring of bat communities along the I-90 project corridor. Illinois Natural History Survey Technical Report 2017(8): 1–15.

21. Rahlin A#, Baker S, Warner JK, Bilger E, Niemiller ML, Beilke S, & Dreslik MJ. Biological monitoring at the North Chicago Wetland Mitigation Site. Illinois Natural History Survey Technical Report 2017(1): 1–21.

2016

20. Engel AS, Niemiller ML, Zigler KS, Fenolio DB, & Kendall KD. Distribution and conservation of cave biodiversity in the Valley and Ridge. Project #A14-0574. Cave Conservancy Foundation. 17 pp.

2015

19. Fenolio DB, Niemiller ML, & McKee AM. Range assessment in Georgia for two threatened groundwater species of the Floridan Aquifer (Cambarus cryptodytes and Eurycea wallacei). Georgia Department of Natural Resources, Wildlife Resources Division, Social Circle, Georgia. 22 pp.

18. Niemiller ML, Zigler KS, & Ober KA. Conservation status of seven Pseudanophthalmus cave beetles and a cave crayfish Orconectes incomptus for possible federal listing. Technical Report. Tennessee Wildlife Resources Agency, Nashville, Tennessee, and U.S. Fish & Wildlife Service, Cookeville, Tennessee. 52 pp.

17. Niemiller ML, & Carter ET. The subterranean fauna of Stamps Cave (TCS No. PU55), Putnam County, Tennessee. Technical Report. Tennessee Parks and Greenways Foundation, Nashville, Tennessee. 16 pp.

2014

16. Armbruster JW, & Niemiller ML. Status survey of Typhlichthys subterraneus in Alabama. Technical Report. Alabama Division of Wildlife and Freshwater Fisheries, Montgomery, Alabama. 31 pp.

2013

15. Niemiller ML, & Fitzpatrick BM. Status, life history, and phylogenetics of amblyopsid cavefishes in Kentucky. Technical Report. Kentucky Department of Fish and Wildlife Resources, Frankfort, Kentucky. 163 pp.

2012

14. Fitzpatrick BM, & Niemiller ML. Status and life history of amblyopsid cavefishes in Kentucky. Pp. 9–15 in Annual Research Highlights, Vol. 5. Kentucky Department of Fish and Wildlife Resources, Frankfort, Kentucky.

2011

13. Fitzpatrick BM, & Niemiller ML. Status, life history, and phylogenetics of amblyopsid cavefishes in Kentucky. Pp. 87 in Annual Research Highlights, Vol. 4. Kentucky Department of Fish and Wildlife Resources, Frankfort, Kentucky.

2010

12. Niemiller ML, Miller BT, & Fitzpatrick BM. Review of the scientific literature and research for the U.S.F.W.S review for potential listing of the Berry Cave salamander (Gyrinophilus gulolineatus). Unpublished Technical Report. U.S. Fish and Wildlife Service. 22 pp.

11. Fitzpatrick BM, & Niemiller ML. Status, life history, and phylogenetics of amblyopsid cavefishes in Kentucky. Pp. 105 in Annual Research Highlights, Vol. 3. Kentucky Department of Fish and Wildlife Resources, Frankfort, Kentucky.

10. Niemiller ML, Miller BT, & Fitzpatrick BM. Status and distribution of the amblyopsid fishes Forbesichthys agassizii and Typhlichthys subterraneus in Tennessee. Unpublished Technical Report. Tennessee Wildlife Resources Agency. 70 pp.

2009

9. Freeman BJ, & Niemiller ML. Species profile for southern cavefish, Typhlichthys subterraneus. Georgia Rare Species Profiles. Georgia Department of Natural Resources, Wildlife Resources Division, Atlanta, Georgia.

8. Fitzpatrick BM, & Niemiller ML. Status, life history, and phylogenetics of amblyopsid cavefishes in Kentucky. Pp. 44 in Annual Research Highlights, Vol. 2. Kentucky Department of Fish and Wildlife Resources, Frankfort, Kentucky.

7. Miller BT, Estabrooks D, Reynolds RG, & Niemiller ML. Status review of the Streamside Salamander (Ambystoma barbouri) in Tennessee. Unpublished Technical Report. Tennessee Wildlife Resources Agency. 42 pp.

2008

6. Reynolds RG, & Niemiller ML. Expedition Report and Recommendations for the Department of Environment and Coastal Resources. Unpublished Technical Report. Ministry of Natural Resources, Turks and Caicos Islands.

5. Fitzpatrick BM, Casper GS, Placyk JS, Niemiller ML, Kirk D, & Burghardt GM. Analysis of the genetic status of populations in the zone of overlap between Thamnophis butleri and Thamnophis radix based on AFLP analysis. Unpublished Technical Report, Wisconsin Department of Natural Resources, Madison. 18 pp.

2007

4. Godwin JC, Jensen JB, Miller BT, & Niemiller ML. Species profile for Tennessee cave salamander, Gyrinophilus palleucus. Georgia Rare Species Profiles. Georgia Department of Natural Resources, Wildlife Resources Division, Atlanta, Georgia.

3. Miller BT, & Niemiller ML. Distribution and phylogenetics of the Tennessee cave salamander complex (Gyrinophilus p. palleucus, G. p. necturoides, and G. gulolineatus). Unpublished Technical Report. Tennessee Wildlife Resources Agency. 45 pp.

2005

2. Miller BT, & Niemiller ML. Distribution, demography, and phylogenetics of the Tennessee cave salamander complex. Unpublished Technical Report. Tennessee Wildlife Resources Agency. 73 pp.

1. Miller BT, Spiess J, & Niemiller ML. Inventory of amphibians and reptiles at Stones River National Battlefield, Rutherford County, Tennessee. Unpublished Technical Report. National Park Service.


Popular Articles

2025

20. Niemiller ML. Cave invaders: introduced species and their impacts on cave ecosystems. NSS News 83(4): 8–9.

19. McCollum S, Helf K, & Niemiller ML. 2025 cave animal of the year – cave shrimp. NSS News 83(4): 6–7.

2024

18. Niemiller ML. Species spotlight: the Berry Cave Salamander. ACCA News 2024(2): 4–6.

17. Niemiller ML, & Christenson K. Clean caving in a white-nose syndrome endemic world. NSS News 8(3): 33–35.

16. Niemiller ML, Helf K, & Fenolio DB. The 2024 cave animal of the year. NSS News 82(3): 8–9.

2022

15. Niemiller ML, Slay ME, & Baker GM. Meet the 2022 USA cave animal of the year: the little brown bat. NSS News 80(4): 5–6.

14. Niemiller ML. Diversity in the darkness: Alabama’s hidden but remarkable life in caves. Alabama Wildlife 86: 19–22.

2021

13. Niemiller ML, & Baker GM. Meet the 2021 USA cave animal of the year. NSS News 79(4): 5.

2020

12. Niemiller ML, Smith W, & Hamed K. Where you can find Green Salamanders and how you can help them. The Tennessee Conservationist 86: 20–25.

2018

11. Engel AS, & Niemiller ML. Landowner’s guide to biological resources and biodiversity in Tennessee caves. In Guide to caves and Sinkholes in Tennessee (Drumm EC, eds). University of Tennessee Extension Institute of Agriculture, University of Tennessee. Publication W453-D. 6 pp.

2017

10. Fenolio D, Niemiller M, Soares D, Gluesenkamp A, Chakrabarty P, & Zhao Y. China’s mysterious cavefishes. Anima Mundi 28: 100–113.

2015

9. Zigler KS, Kuhajda BR, Ayala BN, Niemiller ML, & Engel AS. First observation of southern cavefish (Typhlichthys subterraneus) east of the Cumberland Plateau in northeastern Georgia. Bulletin of the Georgia Speleological Survey 2015: 9–11.

2014

8. Zigler KS, Niemiller ML, & Fenolio DB. Cave biodiversity of the southern Cumberland Plateau of TAG. National Speleological Society Convention Guidebook 2014: 159–163.

7. Niemiller ML, Zigler KS, & Fenolio DB. The obligate cave life of Madison County. SERA Summer Cave Carnival Guidebook.

2012

6. Niemiller ML, Fenolio DB, & Zigler KS. The obligate cave fauna of Georgia. Bulletin of the Georgia Speleological Survey 2012: 6–12.

5. Fenolio D, Bonett R, & Niemiller M. Developing a captive breeding protocol for Georgia’s Blind Salamander, (Haideotriton wallacei) at the Atlanta Botanical Garden. Leaf Litter 1: 40–45.

2011

4. Reynolds, RG, & Niemiller ML. New scales: reptiles invade the Turks and Caicos. Green Pages, Times of the Islands Magazine 94: 30–33.

2010

3. Niemiller ML, & Glorioso BM. The Indo-Pacific Gecko (Hemidactylus garnotii) in Rutherford County, Tennessee. Tennessee Herpetological Society Newsletter.

2005

2. Miller BT, & Niemiller ML. The Tennessee cave salamander complex. Pg. 91–94 in 2005 National Speleological Society Convention Guidebook (Brown JS, & Simon SS eds.). National Speleological Society, Huntsville, AL.

1. Niemiller ML, & Miller BT. Common salamanders of Tennessee caves. Tennessee Caver 2: 12–18.