Journal Articles, Book Chapters, & Proceedings

# Graduate student; ^ Undergraduate student

In press

142. Maxwell EC#, Koltz A^, Stamn J#, Beatty C^, Boudreaux C^, Carbonare J^, Clasper C^, Harden L^, Harris J^, Lawmaster K^, Lee J^, Polk J^, Polk J^, Yeager R^, da Costa Arantes I, Ohmer MEB, & Niemiller ML. Rediscovery of the Tennessee Cave Salamander (Gyrinophilus palleucus) McCrady, 1954 at Shelta Cave, Madison County, Alabama, USA, a former subterranean hotspot cave. Southeastern Naturalist.

141. Niemiller ML, Anderson CL#, Maxwell EC#, Midgette H, Quering C^, Richards M#, Wilson K, & Wirwa N. A noteworthy range extension of the federally endangered Alabama Cave Shrimp (Palaemonias alabamae) south of the Tennessee River in northern Alabama. Southeastern Naturalist.

2025

140. Benito JB#, Ober KA, Philips TK, Orndorff W, & Niemiller ML. Phylogenomics and biogeography of North American trechine cave beetles (Coleoptera: Carabidae) inferred from ultraconserved elements. Evolution 79: 1622–1642. https://doi.org/10.1093/evolut/qpaf103

139. Hart PB, Rincon-Sandoval M, Melendez-Vazquez F, Armbruster JW, Troyer EM, Bierstein OM#, Gough BJ, Betancur-R R, Niemiller ML, & Arcila D. Ancient climate changes and relaxed selection shape cave colonization in North American cavefishes. Proceedings of the Royal Society B, Biological Sciences 292: 20242516. https://doi.org/10.1098/rspb.2024.2516

138. Niemiller ML, Zigler KS, Curtis AN, Trapeni CM^, Slay ME, Culver DC, Hutchins BT, & Niemiller KDK. Out of sight and out of mind? The conservation status of subterranean biodiversity in the United States and Canada. Biodiversity and Conservation 34: 2851–2882. https://doi.org/10.1007/s10531-025-03099-6

137. Niemiller ML, Hart PM, Pirro S, & Arcila D. The complete genome sequences of 12 species of Percopsiformes. Biodiversity Genomes. https://doi.org/10.56179/001c.133607

2024

136. Kennedy KA^, Zigler KS, Cramphorn B#, Harden CW#, Helf K, Lewis JJ, Malabad TE, Milne MA, Niemiller ML, & Stephen CDR. Remarkably low genetic diversity in the widespread cave spider Phanetta subterranea (Araneae, Linyphiidae). Subterranean Biology 50: 105–118. https://doi.org/10.3897/subtbiol.50.135200

135. Cucalón RV#, Corush JB, Niemiller ML, Curtis AN, Hart PB, Kuhajda BR, Thomas MR, Metzke B, Davis MA, & Tan M. 2024. Population genomics and mitochondrial DNA reveal cryptic diversity in North American spring cavefishes (Amblyopsidae, Forbesichthys). Conservation Genetics 25: 1283–1301. https://doi.org/10.1007/s10592-024-01640-8

134. Costley TL#, Hopkins SR, Meng S^, Gajewski Z, & Niemiller ML. Summer cave use by Tricolored Bats declined in response to white-nose syndrome despite persistence in winter hibernacula in the southeastern United States. Journal of Mammalogy 105: 1032–1043. https://doi.org/10.1093/jmammal/gyae064

133. Reynolds RG, Comsa LC#, & Niemiller ML. Status and observations of the critically endangered Turks Island Skink (Spondylurus turksae). Reptiles & Amphibians 31: e22449. https://doi.org/10.17161/randa.v31i1.22449

132. Wilson KL#, Niemiller KDK, & Niemiller ML. The reproductive biology of the northern slimy salamander (Plethodon glutinosus) from a cave in northern Alabama, USA with notes on diet. Herpetological Conservation and Biology 19: 222–235. http://www.herpconbio.org/Volume_19/Issue_2/Wilson_etal_2024.pdf

131. Espinasa L, Gutierrez A^, Hinkle A, & Niemiller ML. A new genus and species of nicoletiid silverfish (Insecta: Zygentoma: Nicoletiidae) from caves of northern Alabama, USA. Subterranean Biology 49: 1–17. https://doi.org/10.3897/subtbiol.49.119986

130. Benito J#, Porter ML, & Niemiller ML. Comparative mitogenomic analysis of subterranean and surface amphipods (Crustacea, Amphipoda) with special reference to the family Crangonyctidae. BMC Genomics 25: 298. https://doi.org/10.1186/s12864-024-10111-w

129. Niemiller ML, Cannizzaro AG#, Sawicki TR, & Culver DC. A new species of Stygobromus Cope, 1872 (Amphipoda: Crangonyctidae) from a hypotelminorheic seepage spring in Washington, D.C., USA. Subterranean Biology 48: 117–146. https://doi.org/10.3897/subtbiol.48.112984

128. Tobin BW, Miller BV, Niemiller ML, & Erhardt AM. Expanding karst groundwater tracing techniques: incorporating population genetic and isotopic data to enhance flow-path characterization. Hydrology 11: 23. https://doi.org/10.3390/hydrology11020023

127. Lunghi E, Niemiller ML, & Bilandzija H. Editorial: Adaptations to subterranean environments. Frontiers in Ecology and Evolution 11: 1354954. https://doi.org/10.3389/fevo.2023.1354954

126. LaFrance BJ, Ray AM, Fisher RN, Grant EHC, Shaffer C, Beamer DA, Spear SF, Pierson TW, Davenport JM, Niemiller ML, Pyron RA, Glorioso BM, Barichivich WJ, Halstead BJ, Roberts KG, & Hossack BR. A dataset of amphibian species in U.S. national parks. Scientific Data 11: 32. https://doi.org/10.1038/s41597-023-02836-2

125. Saccò M, Mammola S, Altermatt F, Alther R, Bolpagni R, Brancelj A, Brankovits D, Fišer C, Gerovasileiou V, Griebler C, Guareschi S, Hose GC, Korbel K, Lictevout E, Malard F, Martínez A, Niemiller ML, Robertson A, Tanalgo KC, Bichuette ME, Borko S, Brad T, Campbell MA, Cardoso P, Celico F, Cooper SJB, Culver D, Di Lorenzo T, Galassi DMP, Guzik MT, Hartland A, Humphreys WF, Ferreira RL, Lunghi E, Nizzoli D, Perina G, Raghavan R, Richards Z, Reboleira ASPS, Rohde MM, Fernández DS, Schmidt SI, van der Heyde M, Weaver L, White NE, Zagmajster M, Hogg I, Gagnon MM, Allentoft ME, & Reinecke R. Groundwater is a globally threatened keystone ecosystem. Global Change Biology 30: e17066. https://doi.org/10.1111/gcb.17066

124. Hart PB, Niemiller ML, Armbruster JW, & Chakrabarty P. Population genomics of a cavefish species complex with implications for conservation. Conservation Genetics 25: 165–177. https://doi.org/10.1007/s10592-023-01562-x

2023

123. Niemiller ML, Zigler KS, Hinkle A, Stephen CDR, Cramphorn B#, Higgs J#, Mann N, Miller BT, Niemiller KDK, Smallwood K, & Hardy J. The Crystal-Wonder Cave System: a new hotspot of subterranean biodiversity in the southern Cumberland Plateau of south-central Tennessee, USA. Diversity 15: 801. https://doi.org/10.3390/d15070801

122. Niemiller ML, Slay ME, Inebnit T, Miller B, Tobin B, Cramphorn B#, Hinkle A, Jones BD, Mann N, Niemiller KDK, & Pitts S. Fern Cave: a hotspot of subterranean biodiversity in the Interior Low Plateau karst region of Alabama in the southeastern United States. Diversity 15: 633. https://doi.org/10.3390/d15050633

121. Mouser JB#, Johnston J, Niemiller ML, & Brewer SK. A fisherman’s tale: an unusual observation of the Ozark cavefish, Troglichthys (=Amblyopsis) rosae (Eigenmann). Southeastern Naturalist 22: N28–N33. https://doi.org/10.1656/058.022.0114

120. Harper LR, Niemiller ML, Benito JB#, Molano-Flores B, & Davis MA. BeeDNA: microfluidic environmental DNA metabarcoding as a tool for connecting plant and pollinator communities. Environmental DNA 5: 191–211. https://doi.org/10.1002/edn3.370

119. Griebler C, Hahn HJ, Mammola S, Niemiller ML, Weaver L, Sacco M, Bichuette ME, & Hose GC. Legal frameworks for the conservation and sustainable management of groundwater ecosystems. Pp. 551–571 in: Groundwater Ecology and Evolution (Malard F, Griebler C, & Retaux S, eds.). Academic Press.

118. Boulton AJ, Bichuette ME, Korbel K, Stoch F, Niemiller ML, Hose GC, & Linke S. Recent concepts and approaches for conserving groundwater biodiversity. Pp. 520–550 in: Groundwater Ecology and Evolution (Malard F, Griebler C, & Retaux S, eds.). Academic Press.

117. Venarsky M, Niemiller ML, Fišer C, Saclier N, & Moldovan OT. Life histories in groundwater organisms. Pp. 439–456 in: Groundwater Ecology and Evolution (Malard F, Griebler C, & Retaux S, eds.). Academic Press.

116. Zagmajster M, Ferreira RL, Humphreys WF, Niemiller ML, & Malard F. Patterns and determinants of richness and composition of the groundwater fauna. Pp. 141–164 in: Groundwater Ecology and Evolution (Malard F, Griebler C, & Retaux S, eds.). Academic Press.

2022

115. Mouser JB#, Brewer SK, Niemiller ML, Mollenhauer R, & Van Den Bussche RA. Lithology and disturbance drive cavefish and cave crayfish occurrence in the Ozark Highlands ecoregion, USA. Scientific Reports 12: 19559. https://doi.org/10.1038/s41598-022-21791-3

114. Grant EHC, Mulder KP, Brand AB, Chambers DB, Wynn A, Capshaw G#, Niemiller ML, Phillips JG, Kuchta SR, & Bell RC. Speciation with gene flow in a narrow endemic West Virginia cave salamander (Gyrinophilus subterraneus). Conservation Genetics 23: 727–744. https://doi.org/10.1007/s10592-022-01445-7

113. Niemiller ML, Hardman R, Thames D, Istvanko D, Davis MA, Ogle C, Niemiller KDK, Dooley KE, & Clark TM. The distribution and conservation status of the green salamander (Aneides aeneus) in Tennessee, USA. Herpetological Conservation and Biology 17: 249–265.

112. Mammola S, Meierhofer M, Borges P, Colado R, Culver D, Deharveng L, Delic T, Di Lorenzo T, Dražina T, Ferreira R, Fiasca B, Fiser C, Galassi D, Garzoli L, Gerovasileiou V, Griebler C, Halse S, Howarth F, Isaia M, Johnson J, Komerički A, Martínez A, Milano F, Moldovan O, Nanni V, Nicolosi G, Niemiller M, Pallarés S, Pavlek M, Piano E, Pipan T, Sánchez Fernández D, Santangeli A, Schmidt S, Wynne J, Zagmajster M, Zakšek V, & Cardoso P. Toward evidence-based conservation of subterranean ecosystems. Biological Reviews 97: 1476–1510. https://doi.org/10.1111/brv.12851

111. Niemiller ML, Davis MA, Tan M, Apodaca JJ, Dooley KE#, Cucalón RV, Benito J#, Niemiller KDK, Harden R, Thames D, & Istvanko D. Mitochondrial DNA and population genomics reveal additional cryptic diversity in the green salamander (subgenus Castaneides) species complex. Frontiers in Conservation Science 3: 890859. https://doi.org/10.3389/fcosc.2022.890859

110. Dooley KE#, Niemiller KDK, Sturm N#, & Niemiller ML. Rediscovery and phylogenetic analysis of the Shelta Cave crayfish (Orconectes sheltae Cooper & Cooper, 1997), a decapod (Decapoda, Cambaridae) endemic to Shelta Cave in northern Alabama, USA. Subterranean Biology 43: 11–31. https://doi.org/10.3897/subtbiol.43.79993

109. Ferreira RL, Bernard E, Júnior FW da C, Piló LB, Calux A, Souza-Silva M, Barlow J, Pompeu PS, Cardoso P, Mammola S, García AM, Jeffery WR, Shear W, Medellín RA, Wynne JJ, Borges PAV, Kamimura Y, Pipan T, Hajna NZ, Sendra A, Peck S, Onac BP, Culver DC, Hoch H, Flot J-F, Stoch F, Pavlek M, Niemiller ML, Manchi S, Deharveng L, Fenolio D, Calaforra J-M, Yager J, Griebler C, Nader FH, Humphreys WF, Hughes AC, Fenton B, Forti P, Sauro F, Veni G, Frumkin A, Gavish-Regev E, Fišer C, Trontelj P, Zagmajster M, Delic T, Galassi DMP, Vaccarelli I, Komnenov M, Gainett G, Tavares V da C, Kováč Ľ, Miller AZ, Yoshizawa K, Lorenzo TD, Moldovan OT, Sánchez-Fernández D, Moutaouakil S, Howarth F, Bilandžija H, Dražina T, Kuharić N, Butorac V, Lienhard C, Cooper SJB, Eme D, Strauss AM, Saccò M, Zhao Y, Williams P, Tian M, Tanalgo K, Woo K-S, Barjakovic M, McCracken GF, Simmons NB, Racey PA, Ford D, Labegalini JA, Colzato N, Pereira MJR, Aguiar LMS, Moratelli R, Preez GD, Pérez-González A, Reboleira ASPS, Gunn J, Cartney AM, Bobrowiec PED, Milko D, Kinuthia W, Fischer E, Meierhofer MB, & Frick WF. Brazilian cave heritage under siege. Science 375: 1238–1239. https://doi.org/10.1126/science.abo1973

108. Gladstone NS#, Niemiller ML, Hutchins B, Schwartz B, Czaja A, Slay ME, & Whelan NV. Subterranean freshwater gastropod biodiversity and conservation in the United States and Mexico. Conservation Biology 36: e13722. https://doi.org/10.1111/cobi.13722

107. Wynne JJ, Niemiller ML, & Chapin KJ. Evolutionary models influencing troglomorphy. Pp. 57–94 in: Cave Life – Drivers of Diversity and Diversification (Wynne JJ, ed.), John Hopkins Press.

106. Niemiller ML, Carter ET, Fenolio DB, Gluesenkamp AG, & Phillips JG. Drivers of subterranean colonization and diversification in cave-dwelling salamanders. Pp. 224–270 in: Cave Life – Drivers of Diversity and Diversification (Wynne JJ, ed.), John Hopkins Press.

105. Ober KA, Niemiller ML, & Philips TK. Cave trechine (Coleoptera: Carabidae) diversity and biogeography in North America. Pp. 192–223 in: Cave Life – Drivers of Diversity and Diversification (Wynne JJ, ed.), John Hopkins Press.

104. Zhao Y, Gluesenkamp AG, Fenolio DB, Soares D, Niemiller ML, Bichuette ME, & Chakrabarty P. Diversity, distribution, and conservation of cavefishes in China. Pp. 271–305 in: Cave Life – Drivers of Diversity and Diversification (Wynne JJ, ed.), John Hopkins Press.

2021

103. Niemiller ML, Carter ET, Gladstone NS#, Niemiller KDK, Hayter LE, Engel AS, Miller BT, & Fitzpatrick BM. The distribution, ecology, life history, and conservation status of the Berry Cave salamander (Gyrinophilus gulolineatus). Herpetological Conservation and Biology 16: 686–703.

102. Wynne JJ, Howarth FG, Mammola S, Ferreira RL, Cardoso P, Di Lorenzo T, Galassi DMP, Medellin RA, Miller BW, Sánchez-Fernández D, Bichuette ME, Biswas J, Boonyanusith C, Borges PAV, Boston PJ, Cheeptham N, Deharveng L, Eme D, Fenolio D, Fišer C, Fišer Z, Gon III S, Goudarzi F, Griebler C, Halse S, Hoch H, Katz AD, Kováč L, Lilley TM, Manchi S, Manenti R, Martínez A, Martinez WE, Meierhofer MB, Miller AZ, Moldovan OT, Niemiller ML, Pipan T, Pellegrini TG, Phillips-Lander CM, Poot C, Racey PA, de Rosário IA, Sendra A, Shear WA, Silva MS, Tian M, Taiti S, Venarsky MP, Pakarati SY, Zagmajster M, & Zhao Y. A roadmap for the conservation of the subterranean biome. Conservation Letters 14: e12834. https://doi.org/10.1111/conl.12834

101. Fitzgerald DB, Smith DR, Culver DC, Feller D, Fong DW, Hajenga J, Niemiller ML, Nolfi DC, Orndorff WD, Douglas B, Maloney KO, & Young JA. Using expert knowledge to support Endangered Species Act decision making for data-deficient species. Conservation Biology 35: 1627–1638. https://doi.org/10.1111/cobi.13694

100. Niemiller KDK, Davis MA, & Niemiller ML. Addressing biodiversity naivety using project-based learning with iNaturalist. Journal for Nature Conservation 64: 126070. https://doi.org/10.1016/j.jnc.2021.126070

99. Niemiller ML, Helf K, & Toomey RS. Mammoth Cave: a hotspot of subterranean biodiversity in the United States. Diversity 13: 373. https://doi.org/10.3390/d13080373

98. Mouser JB#, Brewer SK, Niemiller ML, Mollehauer R, & Van Den Bussche RA. Refining sampling protocols for cavefishes and cave crayfishes to account for environmental variation. Subterranean Biology 39: 79–105. https://doi.org/10.3897/subtbiol.39.64279

97. Gladstone NS#, Pieper EB, Keenan SW, Paterson AT, Slay ME, Dooley K#, Engel AS, & Niemiller ML. Discovery of the Blue Ridge springsnail, Fontigens orolibas, Hubricht, 1957 (Gastropoda: Emmericiidae) in east Tennessee and its conservation implications. Freshwater Mollusk Biology and Conservation 24: 34–42. https://doi.org/10.31931/fmbc-d-20-00007

96. Benito JB#, Porter ML, & Niemiller ML. The mitochondrial genomes of five spring and groundwater amphipods of the family Crangonyctidae (Crustacea: Amphipoda) from eastern North America. Mitochondrial DNA Part B 6: 1662–1667. https://doi.org/10.1080/23802359.2021.1926350

2020

95. Holler Jr C, Mays J, & Niemiller ML. The fauna of caves and other subterranean habitats of North Carolina, USA. Journal of Cave and Karst Studies 82: 221–260. https://doi.org/10.4311/2019LSC0133

94. Soares D, & Niemiller ML. Variation in cephalic neuromasts in surface and cave-dwelling fishes of the family Amblyopsidae (Teleostei: Percopsiformes). Journal of Cave and Karst Studies 82: 198–209. https://doi.org/10.4311/2019LSC0115

93. Boyd SH#, Niemiller KDK, Dooley KE#, Nix J^, & Niemiller ML. Using environmental DNA methods to survey for rare groundwater fauna: detection of an endangered endemic cave crayfish in northern Alabama. PLoS One 15: e0242741. https://doi.org/10.1371/journal.pone.0242741

92. Mammola S, Amorim IR, Bichuette ME, Borges P, Cheeptham N, Cooper SJB, Culver DC, Deharveng L, Eme D, Ferreira RL, Fiser C, Fiser Z, Fong DW, Griebler C, Jeffery WR, Kowalko JE, Jugovic J, Lilley TM, Malard F, Manenti R, Martinez A, Meierhofer MB, Northup DE, Pellegrini TG, Protas M, Niemiller ML, Reboleira AS, Pipan T, Venarsky MP, Wynne JJ, Zagmajster M, & Cardoso P. Fundamental research questions in subterranean biology. Biological Reviews 95: 1855–1872. https://doi.org/10.1111/brv.12642

91. Zigler KS, Niemiller ML, Stephen CDR#, Ayala BN, Milne MA, Gladstone NS#, Engel AS, Jensen JB, & Ozier J. Cave biodiversity of Georgia. Journal of Cave and Karst Studies 82: 125–167. https://doi.org/10.4311/2019LSC0125

90. Baker SJ, Niemiller ML, Stites AJ, Ash KT, Davis MA, Dreslik MJ, & Phillips CA. Evaluation of environmental DNA to detect Sistrurus catenatus and Ophidiomyces ophiodiicola in crayfish burrows. Conservation Genetics Resources 12: 13–15. https://doi.org/10.1007/s12686-018-1053-9

89. DiStefano RJ, Ashley D, Brewer SK, Mouser JB#, & Niemiller ML. Preliminary investigation of the critically imperiled Caney Mountain Cave Crayfish Orconectes stygocaneyi (Decapoda: Cambaridae) (Hobbs III 2001) in Missouri, USA. Freshwater Crayfish 25: 47–57. http://dx.doi.org/10.5869/fc.2020.v25-1.047

88. Hart PB#, Niemiller ML, Burress ED, Armbruster JW, Ludt WB, & Chakrabarty P. Cave-adapted evolution in the North American amblyopsid fishes inferred using phylogenomics and geometric morphometrics. Evolution 74: 936–949. https://doi.org/10.1111/evo.13958

87. Soares D, & Niemiller ML. Extreme adaptation in caves. The Anatomical Record 303: 15–23. https://doi.org/10.1002/ar.24044

2019

86. Gladstone NS#, Perez KE, Pieper EB^, Carter ET, Dooley KE#, Shoobs NF#, Engel AS, & Niemiller ML. A new species of stygobitic snail in the genus Antrorbis Hershler and Thompson, 1990 (Gastropoda, Cochliopidae) from the Appalachian Valley and Ridge of eastern Tennessee, USA. ZooKeys 898: 103–120. https://doi.org/10.3897/zookeys.898.46917

85. Niemiller ML, Inebnit T, Hinkle A, Jones B, Jones M, Lamb J#, Mann N, Miller B, Pinkley J, Pitts S, Sapkota K, & Slay ME. Discovery of a new population of the federally endangered Alabama cave shrimp, Palaemonias alabamae Smalley, 1961, in northern Alabama. Subterranean Biology 32: 43–59. https://doi.org/10.3897/subtbiol.32.38280

84. Wollenberg Valero KC, Marshall JC, Bastiaans E, Caccone A, Camargo A, Morando M, Niemiller ML, Pabijan M, Russello MA, Sinervo B, Werneck JFP, Sites Jr JW, Wiens JJ, & Steinfartz S. Patterns, mechanisms, and genetics of speciation in reptiles and amphibians. Genes 10: 646. https://doi.org/10.3390/genes10090646

83. Mammola S, Cardoso P, Culver DC, Deharveng L, Ferreira RL, Fiser C, Galassi DMP, Griebler C, Halse S, Humphreys WF, Isaia M, Malard F, Martinez A, Moldovan OT, Niemiller ML, Pavlek M, Reboleira ASPS, Souza-Silva M, Teeling EC, Wynne JJ, & Zagmajster M. Scientists’ warning on the conservation of subterranean ecosystems. BioScience 69: 641–650. https://doi.org/10.1093/biosci/biz064

82. Gladstone NS#, Niemiller ML, Pieper EB^, Dooley KE^, & McKinney ML. Morphometrics and phylogeography of the cave-obligate land snail Helicodiscus barri (Gastropoda, Stylommatophora, Helicodiscidae). Subterranean Biology 30: 1–32. https://doi.org/10.3897/subtbiol.30.35321

81. Niemiller ML, & Taylor SJ. Protecting cave life. Pp. 822–829 in: Encyclopedia of Caves, 3rd edition (Culver DC, White WB, & Pipan T, eds.). Elsevier. https://doi.org/10.1016/B978-0-12-814124-3.00099-6

80. Niemiller ML, Taylor SJ, Slay ME, & Hobbs III HH. Biodiversity in the United States and Canada. Pp. 163–176 in: Encyclopedia of Caves, 3rd edition (Culver DC, White WB, & Pipan T, eds.). Elsevier. https://doi.org/10.1016/B978-0-12-814124-3.00018-2

79. Niemiller ML, Bichuette ME, Chakrabarty P, Fenolio DB, Gluesenkamp AG, Soares D, & Zhao Y. Cavefishes. Pp. 227–236 in: Encyclopedia of Caves, 3rd edition (Culver DC, White WB, & Pipan T, eds.). Elsevier. https://doi.org/10.1016/B978-0-12-814124-3.00026-1

78. Goricki S, Niemiller ML, Fenolio DB, & Gluesenkamp AG. Salamanders. Pp. 871–884 in: Encyclopedia of Caves, 3rd edition (Culver DC, White WB, & Pipan T, eds.). Elsevier. https://doi.org/10.1016/B978-0-12-814124-3.00104-7

2018

77. Gladstone NS#, Carter ET, Niemiller KDK, Hayter LE, & Niemiller ML. A new maximum body size record for the Berry Cave salamander (Gyrinophilus gulolineatus) and genus Gyrinophilus (Caudata, Plethodontidae) with a comment on body size in plethodontid salamanders. Subterranean Biology 28: 29–38. https://doi.org/10.3897/subtbiol.28.30506

76. Gladstone NS#, Carter ET, McKinney ML, & Niemiller ML. Status and distribution of cave-obligate land snails in the Appalachians and Interior Low Plateau of the eastern United States. American Malacological Bulletin 36: 62–78. https://doi.org/10.4003/006.036.0107

75. Niemiller ML, Porter ML, Keany J#, Gilbert H, Fong D, Culver DC, Kendall KD, Davis MA, & Taylor SJ. Evaluation of eDNA for groundwater invertebrate detection and monitoring: a case study with endangered Stygobromus (Amphipoda: Crangonyctidae). Conservation Genetics Resources 10: 247–257. https://doi.org/10.1007/s12686-017-0785-2

74. Niemiller ML, Bichuette E, & Taylor SJ. Conservation of cave fauna in Europe and the Americas. Pp. 451–478 in: Ecological Studies: Cave Ecology (Moldovan OT, Kovac L, & Halse S, eds). Springer. http://dx.doi.org/10.1007/978-3-319-98852-8_22

2017

73. Soares D, Adams R#, Hammond S, Slay ME, Fenolio DB, & Niemiller ML. Evolution of coprophagy and nutrient absorption in a cave salamander. Subterranean Biology 24: 1–9. https://doi.org/10.3897/subtbiol.24.15013

72. Nelson SK, Niemiller ML, & Fitzpatrick BM. Co-occurrence and hybridization between Necturus maculosus and an undescribed Necturus species in the Southern Appalachians. Journal of Herpetology 51: 559–566. https://doi.org/10.1670/17-006

71. Niemiller ML, Zigler KS, Ober KA, Carter ET#, Engel AS, Moni G, Philips TK, & Stephen CDR#. Rediscovery and conservation status of six short-range endemic Pseudanophthalmus cave beetles (Carabidae: Trechini). Insect Conservation and Diversity 10: 495–501. https://doi.org/10.1111/icad.12263

70. Fenolio DB, Niemiller ML, Gluesenkamp AG, McKee AM, & Taylor SJ. New distributional records of the stygobiotic crayfish Cambarus cryptodytes (Decapoda: Cambaridae) in the Floridan aquifer system of southwestern Georgia. Southeastern Naturalist 16: 163–181. https://doi.org/10.1656/058.016.0205

69. Bagley R#, Sousa V, Niemiller ML, & Linnen C. History, geography, and host use shape genome-wide patterns of genetic variation in the redheaded pine sawfly (Neodiprion lecontei). Molecular Ecology 26: 1022–1044. https://doi.org/10.1111/mec.13972

68. Engel AE, Paterson AT, & Niemiller ML. Biogeography of microbes from caves in the Interior Low Plateau and Appalachian karst regions. Pp. 410 in: Proceedings of the 17th International Congress of Speleology, Volume 1 (Moore K, & White S). Australian Speleological Federation & Speleo2017.

67. Keenan SW, Paterson AT, Niemiller ML, Slay ME, Clark SA, & Engel AS. Observations of the first stygobiont snail (Hydrobiidae, Fontigens sp.) in Tennessee. Pp. 91–94 in: Proceedings of the 17th International Congress of Speleology, Volume 1 (Moore K, & White S). Australian Speleological Federation & Speleo2017.

66. Engel AS, Niemiller ML, Zigler KS, Stephen CDR, Carter ET, Paterson AT, Keenan SW, & Taylor SJ. Invertebrate and vertebrate cave fauna records for the Appalachian Valley and Ridge. Pp. 82–86 in: Proceedings of the 17th International Congress of Speleology, Volume 1 (Moore K, & White S). Australian Speleological Federation & Speleo2017.

2016

65. Niemiller ML, Zigler KS, Hart PB#, Kuhajda BR, Ayala BN^, Armbruster JW, & Engel AS. First record of a stygobiotic fish (Percopsiformes: Amblyopsidae: Typhlichthys) from the Appalachians karst region in the eastern United States. Subterranean Biology 20: 39–50. https://doi.org/10.3897/subtbiol.20.9693

64. Armbruster JW, Niemiller ML, & Hart PB#. Morphological evolution of the cave-, spring- and swampfishes of the family Amblyopsidae (Percopsiformes). Copeia 104: 763–777. https://doi.org/10.1643/CI-15-339

63. Christman MC, Doctor DH, Niemiller ML, Weary DJ, Young JA, Zigler KS, & Culver DC. Predicting the occurrence of cave-inhabiting fauna based on features of the Earth surface environment. PLoS One 11: e0160408. https://doi.org/10.1371/journal.pone.0160408

62. Reynolds RG, Collar DC, Pasachnik SA, Niemiller ML, Puente-Rolon AR, & Revell LJ. Ecological specialization and morphological diversification in Greater Antillean boas. Evolution 70: 1882–1895. https://doi.org/10.1111/evo.12987

61. Niemiller ML, Zigler KS, Stephen CDR#, Carter ET#, Paterson AT, Taylor SJ, & Engel AS. Vertebrate fauna in caves of eastern Tennessee within the Appalachians karst region, USA. Journal of Cave and Karst Studies 78: 1–24. https://doi.org/10.4311/2015lsc0109

60. Niemiller ML, Glorioso BM, Fenolio DB, Reynolds RG, Taylor SJ, & Miller BT. Growth, survival, longevity, and population size of the Big Mouth Cave salamander (Gyrinophilus palleucus necturoides) from the type locality in Grundy Co., Tennessee, USA. Copeia 104: 35–41. https://doi.org/10.1643/OT-14-197

59. Taylor SJ, & Niemiller ML. Biogeography and conservation of Bactrurus groundwater amphipods (Crangonyctidae) in the central and eastern United States. Subterranean Biology 17: 1–29. https://doi.org/10.3897/subtbiol.17.7298

58. Soares D, Niemiller ML, & Higgs DM. Hearing in cavefishes. Pp. 187–195 in: Fish Hearing and Bioacoustics – An Anthology in Honor of Arthur N. Popper and Richard R. Fay (Sisneros JA, ed). Springer. https://doi.org/10.1007/978-3-319-21059-9_9

2015

57. Taylor SJ, Krejca JK, Niemiller ML, Dreslik MJ, & Phillips CA. Life history and demographic differences between cave and surface populations of the western slimy salamander Plethodon albagula (Caudata: Plethodontidae) in central Texas. Herpetological Conservation and Biology 10: 740–752.

56. Niemiller ML*, & Soares D*. Cave environments. Pp. 161–191 in: Extremophile Fishes – Ecology and Evolution of Teleosts in Extreme Environments (Riesch R, Plath M, & Tobler M, eds). Springer. https://doi.org/10.1007/978-3-319-13362-1_8

2014

55. Thompson S, Muzinic L, Niemiller ML, & Voss SR. Probability of regenerating a normal limb after bite injury in the Mexican axolotl (Ambystoma mexicanum). Regeneration 1: 27–32. https://doi.org/10.1002/reg2.17

54. Kendall KD, Niemiller ML, Dittrich-Reed D, & Schussler EE. Helping graduate teaching assistants in biology use student evaluations as professional development. The American Biology Teacher 76: 584–588. https://doi.org/10.1525/abt.2014.76.9.3

53. Fenolio DB, Niemiller ML, Bonett RM, Graening GO, Collier BA, & Stout JF. Life history, demography, and the influence of cave-roosting bats on a population of the grotto salamander (Eurycea spelaea) from the Ozark Plateaus of Oklahoma (Caudata: Plethodontidae). Herpetological Conservation and Biology 9: 394–405.

52. Espinasa L, Espinasa M, Fenolio DB, Slay ME, & Niemiller ML. Distribution and conservation status of Speleonycta ozarkensis (Insecta: Zygentoma: Nicoletiidae) from caves of the Ozark Highlands of Arkansas and Oklahoma, USA. Subterranean Biology 14: 51–62. https://doi.org/10.3897/subtbiol.14.8275

51. Chakrabarty P, Prejean JA, & Niemiller ML. The Hoosier cavefish, a new and endangered species (Amblyopsidae, Amblyopsis) from the caves of southern Indiana. ZooKeys 412: 41–57. https://doi.org/10.3897/zookeys.412.7245

50. Soares D, Niemiller ML, & Higgs D. Hearing and acoustic communication in cavefishes. Ambient Science 1: 1–6.

49. Kennan SW, Niemiller ML, & Williams BW. Observations of Cambarus bartonii cavatus (Decapoda: Cambaridae) and ectosymbiotic branchiobdellidans (Annelida: Clitellata) in Cruze Cave, Knox County, Tennessee, USA. Speleobiology Notes 6: 55–61.

48. Fenolio DB, Niemiller ML, & Martinez B. Observations of reproduction in captivity by the Dougherty Plain Cave Crayfish, Cambarus cryptodytes (Decapoda: Astacoidea: Cambaridae). Speleobiology Notes 6: 14–26.

47. Reynolds RG, Niemiller ML, & Revell LJ. Toward a Tree-of-Life for the boas and pythons: multilocus species-level phylogeny with unprecedented taxon sampling. Molecular Phylogenetics and Evolution 71: 201–213. https://doi.org/10.1016/j.ympev.2013.11.011

46. Niemiller ML, & Reeves WC. Predation by the stygophilic crayfish Cambarus tenebrosus on the salamander Pseudotriton ruber within a cave in Cannon County, Tennessee, USA. Speleobiology Notes 6: 8–13.

2013

45. Barun A, Niemiller ML, Fitzpatrick BM, Fordyce JA, & Simberloff D. Can genetic data confirm or refute historical records: the island invasion of the small Indian mongoose (Herpestes auropunctatus). Biological Invasions 15: 2243–2251. https://doi.org/10.1007/s10530-013-0447-6

44. Fenolio DB, Niemiller ML, Levy M, & Martinez B. Conservation status of the Georgia blind salamander (Eurycea wallacei) from the Floridan Aquifer of Florida and Georgia. Reptiles and Amphibians 20: 97–111. https://doi.org/10.17161/randa.v20i3.13945      

43. Kendall KD, Niemiller ML, Dittrich-Reed D, Chick LD, Wilmoth L, Milt A, Burt M, Lopes N, Cantwell L, Rubio L, Allison A, & Schussler EE. Departments can develop teaching identities of graduate students. CBE – Life Sciences Education 12: 316–317. https://doi.org/10.1187/cbe.13-03-0066

42. McAllister CT, Bursey CR, Fenolio DB, & Niemiller ML. Bothriocephalus sp. (Cestoidea: Bothriocephalidea: Bothriocephalidae) from the Georgia blind salamander, Eurycea wallacei (Caudata: Plethodontidae), in Georgia, USA: first definitive report of a parasite from this host. Comparative Parasitology 80: 308–311. https://doi.org/10.1654/4615.1

41. Niemiller ML, Graening GO, Fenolio DB, Godwin JC, Cooley JR, Pearson WR, Near TJ, & Fitzpatrick BM. Doomed before they are described? The need for conservation assessments of cryptic species complexes using an amblyopsid cavefish (Amblyopsidae: Typhlichthys) as a case study. Biodiversity and Conservation 22: 1799–1820. https://doi.org/10.1007/s10531-013-0514-4

40. Reynolds RG, Niemiller ML, Hedges SB, Dornburg A, & Revell LJ. Molecular phylogeny and historical biogeography of West Indian boid snakes (Chilabothrus). Molecular Phylogenetics and Evolution 68: 461–470. https://doi.org/10.1016/j.ympev.2013.04.029

39. Niemiller ML, & Zigler KS. Patterns of cave biodiversity and endemism in the Appalachians and Interior Plateau of Tennessee USA. PLoS One 8: e64177. https://doi.org/10.1371/journal.pone.0064177

38. Fenolio DB, Niemiller ML, Soares D, Slay ME, Harris KA, & Harris N. Subterranean reproduction of the ringed crayfish, Orconectes neglectus Faxon 1885 (Astacoidea: Cambaridae) within a cave in the Ozark Highlands of Oklahoma, USA. Speleobiology Notes 5: 43–46.

37. Fenolio DB, McAllister CT, Niemiller ML, Soares D, & Cooley J. An extreme case of trematode parasitic infection of a larval Ozark blind cave salamander, Eurycea spelaea (Caudata: Plethodontidae) from the Ozark Highlands of Missouri, USA. Speleobiology Notes 5: 34–37.

36. Fenolio DB, Zhao Y, Niemiller ML, & Stout JF. In situ observations of seven enigmatic cave loaches and one cave barbel from Guangxi, China, with notes on conservation status. Speleobiology Notes 5: 19–33.

35. Fenolio DB, Niemiller ML, Soares D, Slay ME, Stark RC, & Hensley SL. A maximum size and maximum abundance record of the Delaware County cave crayfish, Cambarus subterraneus Hobbs 1993 (Astacoidea: Cambaridae), from Twin Cave, Delaware County, Oklahoma, USA. Speleobiology Notes 5: 9–13.

34. Soares D, & Niemiller ML. Sensory adaptations of cavefishes to cave environments. BioScience 63: 274–283. https://doi.org/10.1525/bio.2013.63.4.7

33. Niemiller ML, McCandless JR, Reynolds RG, Caddle J, Tillquist CR, Near TJ, Pearson WD, & Fitzpatrick BM. Effects of climatic and geological processes during the Pleistocene on the evolutionary history of the northern cavefish, Amblyopsis spelaea (Teleostei: Amblyopsidae). Evolution 67: 1011–1025. https://doi.org/10.1111/evo.12017

32. Niemiller ML, Higgs DM, & Soares D. Evidence for hearing loss in amblyopsid cavefishes. Biology Letters 9: 20130104. https://doi.org/10.1098/rsbl.2013.0104

31. Niemiller ML, Fitzpatrick BM, Shah P, Schmitz L, & Near TJ. Evidence for repeated loss of selective constraint in rhodopsin of amblyopsid cavefishes. Evolution 67: 732–748. https://doi.org/10.1111/j.1558-5646.2012.01822.x

2012

30. Reynolds RG, Niemiller ML, & Fitzpatrick BM. Genetic analysis of an endemic archipelagic lizard reveals sympatric cryptic lineages and taxonomic discordance. Conservation Genetics 13: 953–963. https://doi.org/10.1007/s10592-012-0344-z

29. Miller BT, & Niemiller ML. Gyrinophilus palleucus. Catalogue of American Amphibians and Reptiles 884: 1–7.

28. Niemiller ML, Near TJ, & Fitzpatrick BM. Delimiting species using multilocus data: diagnosing cryptic diversity in the southern cavefish Typhlichthys subterraneus (Teleostei: Amblyopsidae). Evolution 66: 846–866. https://doi.org/10.1111/j.1558-5646.2011.01480.x

287 Fitzpatrick BM, Fordyce JA, Niemiller ML, & Reynolds RG. What can DNA tell us about biological invasions? Biological Invasions 14: 245–253. https://doi.org/10.1007/s10530-011-0064-1

26. Goricki A, Niemiller ML, & Fenolio DB. Salamanders. Pp. 665–676 in: Encyclopedia of Caves, 2nd Edition (White WH, & Culver DC, eds.). Elsevier. https://doi.org/10.1016/b978-0-12-383832-2.00098-0

2011

25. Niemiller ML, Reynolds RG, Glorioso BM, Spiess J, & Miller BT. Herpetofauna of the cedar glades and associated habitats of the Inner Central Basin of middle Tennessee. Herpetological Conservation and Biology 6: 135–149.

24. Reynolds RG, Niemiller ML, & Pasachnik SA. Distribution of the Pygmy Salamander (Desmognathus wrighti) in Tennessee. Journal of the Tennessee Academy of Science 86: 53–55.

2010

23. Glorioso BM, Niemiller ML, & Cobb VA. Feeding activity of Amphiuma tridactylum at Reelfoot Lake, Tennessee. Journal of the Tennessee Academy of Science 85: 87–90.

22. Reynolds RG, & Niemiller ML. 2010. Island invaders: introduced reptiles and amphibians of the Turks and Caicos Islands. Reptiles and Amphibians 17: 117–121.

21. Niemiller ML, Osbourn MS, Fenolio DB, Pauley TK, Miller BT, & Holsinger JR. Conservation status and habitat use of the West Virginia Spring Salamander (Gyrinophilus subterraneus) and Spring Salamander (G. porphyriticus) in General Davis Cave, Greenbrier Co., West Virginia. Herpetological Conservation and Biology 5: 32–43.

20. Niemiller ML, & Miller BT. Gyrinophilus gulolineatus. Catalogue of American Amphibians and Reptiles. 862.1.

19. Niemiller ML, Nosil P, & Fitzpatrick BM. Corrigendum. Recent divergence-with-gene-flow in Tennessee cave salamanders (Plethodontidae: Gyrinophilus) inferred from gene genealogies. Molecular Ecology 19: 1513–1514. https://doi.org/10.1111/j.1365-294x.2010.04564.x

18. Graening GO, Fenolio DB, Niemiller ML, Brown AV, & Beard JB. The 30-year recovery effort for the Ozark cavefish (Amblyopsis rosae): analysis of current distribution, population trends, and conservation status of this threatened species. Environmental Biology of Fishes 87: 55–88. https://doi.org/10.1007/s10641-009-9568-2

17. Niemiller ML, & Poulson TL. Subterranean fishes of North America: Amblyopsidae. Pp. 169–280 in: The Biology of Subterranean Fishes (Trajano E, Bichuette ME, & Kapoor BG, eds). Science Publishers, Enfield, New Hampshire, USA. https://doi.org/10.1201/ebk1578086702

2009

16. Niemiller ML, Glorioso BM, Nicholas C, Phillips J, Rader J, Reed E, Sykes KL, Todd J, Wyckoff GR, Young EL, & Miller BT. Notes on the reproduction of the streamside salamander, Ambystoma barbouri, from Rutherford County, Tennessee. Southeastern Naturalist 8: 37–44. https://doi.org/10.1656/058.008.0104

15. Niemiller ML, Fenolio D, Graening GO, & Miller BT. Observations on oviposition and reproduction of the cave salamander, Eurycea lucifuga (Caudata: Plethodontidae), from Arkansas and Tennessee. Speleobiology Notes 1: 17–19.

14. Niemiller ML, Miller BT, & Fitzpatrick BM. Systematics and evolutionary history of subterranean Gyrinophilus salamanders. Proceedings of the 15th International Congress of Speleology, Kerrville, Texas 15: 242–248.

13. Niemiller ML, & Miller BT. A survey of the cave-associated amphibians of the eastern United States with an emphasis on salamanders. Proceedings of the 15th International Congress of Speleology, Kerrville, Texas 15: 249–256.

2008

12. Miller BT, Niemiller ML, & Reynolds RG. Observations on egg-laying behavior and interactions among attend female red salamanders (Pseudotriton ruber) with comments on the use of caves by this species. Herpetological Conservation and Biology 3: 203–210.

11. Fitzpatrick BM, Placyk Jr JS, Niemiller ML, Casper GS, & Burghardt GM. Distinctiveness in the face of gene flow: hybridization between specialist and generalist gartersnakes. Molecular Ecology 17: 4107–4117. https://doi.org/10.1111/j.1365-294x.2008.03885.x

10. Niemiller ML, Fitzpatrick BM, & Miller BT. Recent divergence-with-gene-flow in Tennessee cave salamanders (Plethodontidae: Gyrinophilus) inferred from gene genealogies. Molecular Ecology 17: 2258–2275. https://doi.org/10.1111/j.1365-294x.2008.03750.x

9. Miller BT, & Niemiller ML. Distribution and relative abundance of Tennessee cave salamanders (Gyrinophilus palleucus and G. gulolineatus) with an emphasis on Tennessee populations. Herpetological Conservation and Biology 3: 1–20.

8. Niemiller ML, & Fitzpatrick BM. Phylogenetics of the southern cavefish, Typhlichthys subterraneus: implications for conservation and management. Proceedings of the 18th National Cave and Karst Management Symposium 18: 79–88.

2007

7. McAllister CT, Bursey CR, Niemiller ML, & Miller BT. A noteworthy infection of Clinostomum complanatum (Digenea: Clinostomidae) in a cave salamander, Eurycea lucifuga (Caudata: Plethodontidae), from northcentral Tennessee. Texas Journal of Science 59: 321–326.

6. Niemiller ML, & Miller BT. Subterranean reproduction of the southern two-lined salamander (Eurycea cirrigera) from Short Mountain, Tennessee. Herpetological Conservation and Biology 2: 106–112.

2006

5. Niemiller ML, Glorioso BM, Nicholas C, Phillips J, Rader J, Reed E, Sykes KL, Todd J, Wyckoff GR, Young EL, & Miller BT. Status and distribution of the streamside salamander (Ambystoma barbouri) in middle Tennessee. American Midland Naturalist 156: 393–399. https://doi.org/10.1674/0003-0031(2006)156[394:sadots]2.0.co;2

4. Glorioso BM, & Niemiller ML. Using deep-water crawfish nets to capture aquatic turtles. Herpetological Review 37: 185–187.

3. Niemiller ML, Glorioso BM, & Miller BT. An eastern and subterranean maximum size record of the banded sculpin, Cottus carolinae, from Tennessee. Journal of the Tennessee Academy of Science 81: 41–43.

2005

2. Koczaja C, McCall L, Fitch E, Glorioso B, Hanna C, Kyzar J, Niemiller M, Spiess J, Tolley A, Wyckoff R, & Mullen D. Size-specific habitat segregation and intraspecific interactions in banded sculpin (Cottus carolinae). Southeastern Naturalist 4: 207–218. https://doi.org/10.1656/1528-7092(2005)004[0207:shsaii]2.0.co;2

1. Niemiller ML. The herpetofauna of the upper Duck River watershed in Coffee County, Tennessee. Journal of the Tennessee Academy of Science 80: 6–12.